References

Alcock, J. (Ed.) (1998). ‘Animal Behavior: an Evolutionary Approach.’ (Sinauer: Sunderland, MA.)

Arck, P. C., Merali, F. S., Stanisz, A. M., Stead, R. H., Chaouat, G., and Manuel, J. (1995). Stress-induced murine abortion associated with substance P-dependent alteration in cytokines in maternal uterine decidua. Biology of Reproduction 53, 814–819.
| CrossRef | CAS |

Balmford, A., Beresford, J., Green, J., Naidoo, R., Walpole, M., and Manica, A. (2009). A global perspective on trends in nature-based tourism. PLoS Biology 7, e1000144.
| CrossRef |

Barja, I. (2005). Winter distribution of European pine marten (Martes martes) scats in a protected area of Galicia, Spain. Mammalia 69, 435–438.
| CrossRef |

Barja, I., and Bárcena, F. (2005). Distribución y abundancia de gato montés (Felis silvestris) en el Parque Natural Os Montes do Invernadeiro (Galicia, NO de España): factores de hábitat implicados y relación con la presencia de zorro y marta. Galemys 17, 29–40.

Barja, I., Silván, G., Rosellini, S., Piñeiro, A., González-Gil, A., Camacho, L., and Illera, J. C. (2007). Stress physiological responses to tourist pressure in a wild population of European pine marten. The Journal of Steroid Biochemistry and Molecular Biology 104, 136–142.
| CrossRef | CAS |

Barja, I., Silván, G., Rosellini, S., Piñeiro, A., Illera, M. J., and Illera, J. C. (2008a). Quantification of sexual steroid hormones in faeces of Iberian wolf (Canis lupus signatus): a non-invasive sex typing method. Reproduction in Domestic Animals 43, 701–707.
| CrossRef | CAS |

Barja, I., Silván, G., and Illera, J. C. (2008b). Relationships between sexual and stress hormone levels in feces and marking behavior in a wild population of Iberian wolves (Canis lupus signatus). Journal of Chemical Ecology 34, 697–701.
| CrossRef | CAS |

Barja, I., Silván, G., Martínez-Fernández, L., and Illera, J. C. (2011). Physiological stress responses, faecal marking behaviour, and reproduction in wild European pine martens (Martes martes). Journal of Chemical Ecology 37, 253–259.
| CrossRef | CAS |

Belkhir, K., Borsout, P., Goudet, J., Chikhi, L., and Bonhomme, F. (Eds) (1996–2004). ‘Genetix, Logiciel sous WindowsTM pour la Génétique des Populations,’ (Laboratoire Génome et Population, Université de Montpellier: Montepellier, France.)

Benton, T. G., Vickery, J. A., and Wilson, J. D. (2003). Farmland biodiversity: is habitat heterogeneity the key? Trends in Ecology & Evolution 18, 182–188.
| CrossRef |

Bonier, F., Quigley, H., and Austad, S. N. (2004). A technique for non-invasively detecting stress response in cougars. Wildlife Society Bulletin 32, 711–717.
| CrossRef |

Brown, J. L., and Wildt, D. E. (1997). Assessing reproductive status in wild felids by non-invasive faecal steroid monitoring. International Zoo Yearbook 35, 173–191.
| CrossRef |

Brown, J. L., Wasser, S. K., Wildt, D. E., and Graham, L. H. (1994). Comparative aspects of steroid hormone metabolism and ovarian activity in felids, measured non invasively in feces. Biology of Reproduction 51, 776–786.
| CrossRef | CAS |

Brown, J. L., Wildt, D. E., Wielebnowski, N., Goodrowe, K. L., Graham, L. H., Wells, S., and Howard, J. G. (1996). Reproductive activity in captive female cheetahs (Acinonyx jubatus) assessed by faecal steroids. Journal of Reproduction and Fertility 106, 337–346.
| CrossRef | CAS |

Burnham, K. P., and Anderson, D. R. (2002). ‘Model Selection and Multimodel Inference. A Practical Information-theoretic Approach.’ 2nd edn. (Springer: New York.)

Busch, D. S., and Hayward, L. S. (2009). Stress in a conservation context: a discussion of glucocorticoid actions and how levels change with conservation-relevant variables. Biological Conservation 142, 2844–2853.
| CrossRef |

Carlstead, K., and Shepherdson, D. (1994). Effects of environmental enrichment on reproduction. Zoo Biology 13, 447–458.
| CrossRef |

Corbett, L. K. (1979). Feeding ecology and social organization of wildcats (Felis silvestris) and domestics cats (Felis catus) in Scotland. Ph.D. Thesis, University of Aberdeen, Scotland.

Creel, S., Fox, J. E., Hardy, A., Sands, J., Garrott, B., and Peterson, R. O. (2002). Snowmobile activity and glucocorticoid stress responses in wolves and elk. Conservation Biology 16, 809–814.
| CrossRef |

Dehnhard, M., Clauss, M., Lechner-Doll, M., Meyer, H. H. D., and Palme, R. (2001). Noninvasive monitoring of adrenocortical activity in roe deer (Capreolus capreolus) by measurement of faecal cortisol metabolites. General and Comparative Endocrinology 123, 111–120.
| CrossRef | CAS |

Eggermann, J. (2009). The impact of habitat fragmentation by anthropogenic infraestructures on wolves (Canis lupus). Ph.D. Thesis, University of Bochum, Germany.

Gagliuso, R. A. (1992). Habitat alteration and human disturbance: their impact on cougar habitat utilization in Southwest Oregon. Ph.D. Thesis, University of Oregon, Corvallis, OR, USA.

Graham, L. H., and Brown, J. L. (1996). Cortisol metabolism in the domestic cat and implications for non-invasive monitoring of adrenocortical function in endangered felids. Zoo Biology 15, 71–82.
| CrossRef | CAS |

Graham, L. H., Goodrowe, K. L., Raeside, J. I., and Liptrap, R. M. (1995). Non-invasive monitoring of ovarian function in several felid species by measurement of faecal estradiol-17β and progestins. Zoo Biology 14, 223–237.
| CrossRef | CAS |

Greenberg, N., Carr, J. A., and Summers, C. H. (2002). Causes and consequences of stress. Integrative and Comparative Biology 42, 508–516.
| CrossRef |

IUCN (2011). ‘IUCN Red List of Threatened Species. Version 2011.2.’ Available at www.iucnredlist.org [verified 27 November 2011].

Jerosch, S., Götz, M., Klar, N., and Roth, M. (2010). Characteristics of diurnal resting sites of the endangered European wildcat (Felis silvestris silvestris): Implications for its conservation. Journal for Nature Conservation 18, 45–54.
| CrossRef |

Jurke, M. H., Czekala, N. M., Lindburg, D. G., and Millard, S. E. (1997). Faecal corticoid metabolite measurement in the cheetah (Acinonyx jubatus). Zoo Biology 16, 133–147.
| CrossRef | CAS |

Kerley, L. L., Goodrich, J. M., Miquelle, D. G., Smirnov, E. N., Quigley, H. B., and Hornocker, M. G. (2002). Effects of roads and human disturbance on Amur tigers. Conservation Biology 16, 97–108.
| CrossRef |

Kitchener, A. (Ed.) (1995). ‘The Wildcat.’ (The Mammal Society: London.)

Klar, N., Fernández, N., Kramer-Schadt, S., Herrmann, M., Trinzenf, M., Büttner, I., and Niemitz, C. (2008). Habitat selection models for European wildcat conservation. Biological Conservation 141, 308–319.
| CrossRef |

Liu, J., Chen, Y., Guo, L., Gu, B., Liu, H., Hou, A., Liu, X., Sun, L., and Liu, D. (2006). Stereotypic behavior and faecal cortisol level in captive giant pandas in relation to environmental enrichment. Zoo Biology 25, 445–459.
| CrossRef | CAS |

Mellen, J. (Ed.) (2002). ‘Husbandry Standards for Keeping Small Cats in Captivity.’ (American Zoo and Aquarium Association: Silver Spring, MD.)

Millspaugh, J. J., and Washburn, B. E. (2003). Within-sample variation of fecal glucocorticoid measurements. General and Comparative Endocrinology 132, 21–26.
| CrossRef | CAS |

Moberg, G. P. (Ed.) (1985). ‘Animal Stress.’ (American Physiological Society: Bethesda, MD.)

Monfort, S. L., Mashburn, K. L., Brewer, B. A., and Creel, S. R. (1998). Evaluating adrenal activity in African wild dogs (Lycaon pictus) by faecal corticosteroid analysis. Journal of Zoo and Wildlife Medicine 29, 129–133.
| CAS |

Montanha, J. C., Silva, S. L., and Boere, V. (2009). Comparison of salivary cortisol concentrations in jaguars kept in captivity with differences in exposure to the public. Ciência Rural 39, 1745–1751.
| CrossRef | CAS |

Monterroso, P., Brito, J. C., Ferreras, P., and Alves, P. C. (2009). Spatial ecology of the European wildcat in a Mediterranean ecosystem: dealing with small radio-tracking datasets in species conservation. Journal of Zoology 279, 27–35.
| CrossRef |

Möstl, E., and Palme, R. (2002). Hormones as indicators of stress. Domestic Animal Endocrinology 23, 67–74.
| CrossRef |

Müllner, A., Linsenmair, K. E., and Wikelski, M. (2004). Exposure to ecotourism reduces survival and affects stress response in hoatzin chicks (Opisthocomus hoazin). Biological Conservation 118, 549–558.
| CrossRef |

Nepomnaschy, P. A., Welch, K. B., McConnell, D. S., Low, B. S., Strassman, B. I., and England, B. G. (2006). Cortisol levels and very early pregnancy loss in humans. Proceedings of the National Academy of Sciences, USA 103, 3938–3942.
| CrossRef | CAS |

Ngoprasert, D., Lynam, A. J., and Gale, G. A. (2007). Human disturbance affects habitat use and behaviour of Asiatic leopard Panthera pardus in Kaeng Krachan National Park, Thailand. Oryx 41, 343–351.
| CrossRef |

Nowell, K., and Jackson, P. (Eds) (1996). ‘The Wild Cats: Status Survey and Conservation Action Plan.’ (IUCN/Cat Specialist Group: Gland, Switzerland.)

Oliveira, R., Godinho, R., Randi, E., and Alves, P. C. (2008). Hybridization versus conservation: are domestic cats threatening the genetic integrity of wildcats (Felis silvestris silvestris) in Iberian Peninsula? Philosophical Transactions of the Royal Society of London. B 363, 2953–2961.
| CrossRef |

Parmesan, C. (2006). Ecological and evolutionary responses to recent climate change. Annual Review of Ecology Evolution and Systematics 37, 637–669.
| CrossRef |

Pierce, B. N., Hemsworth, P. H., Rivalland, E. T. A., Wagenmaker, E. R., Morrissey, A. D., Papargiris, M. M., Clarke, I. J., Karsch, F. J., Turner, A. I., and Tilbrook, A. J. (2008). Psychosocial stress suppresses attractivity, proceptivity and pulsatile LH secretion in the ewe. Hormones and Behavior 54, 424–434.
| CrossRef | CAS |

Pulgar, I. (Ed.) (2004). ‘Guía da flora do Parque Natural Montes do Invernadeiro.’ (Consellería de Medio Ambiente, Xunta de Galicia: Ourense, Spain.)

Randi, E., Pierpaoli, M., Beaumont, M., Ragni, B., and Sforzi, A. (2001). Genetic identification of wild and domestic cats (Felis silvestris) and their hybrids using bayesian clustering methods. Molecular Biology and Evolution 18, 1679–1693.
| CrossRef | CAS |

Sambrook, E., Fritsch, F., and Maniatis, T. (Eds) (1989). ‘Molecular Cloning.’ (Cold Spring Harbour Press: New York.)

Sapolsky, R. M. (1986). Stress-induced elevation of testosterone concentrations in high-ranking baboons: role of catecholamines. Endocrinology 118, 1630–1635.
| CrossRef | CAS |

Sapolsky, R. M. (1992). Neuroendocrinology of the stress response. In ‘Behavioral Endocrinology’. (Eds J. B. Becker, S. M. Breedlove and D. Crews.) pp. 287–324. (MIT Press: Cambridge, UK.)

Sapolsky, R. M., Romero, L. M., and Munck, A. U. (2000). How do glucocorticoids influence stress responses? Integrating permissive, suppressive, stimulatory, and preparative actions. Endocrine Reviews 21, 55–89.
| CrossRef | CAS |

Schwarzenberger, F., Walzer, C., Weiss, R., Pucher, H. E., Leus, K., Bircher, P., and Moran, J. F. (2000). Faecal cortisol metabolites in response to management practices and disturbance in different herbivores and carnivores (Ceratotherium simum, Diceros bicornis, Elephas maximus, Loxodonta africana, Okapia johnstoni, Rupicapra rupicapra and Lutra lutra). In ‘Proceedings of the European Association of Zoo and Wildlife Veterinarians (EAZWV), Third Scientific Meeting’. (Ed. P. Zwart.) pp. 1–4. (Paris Zoo: Paris.)

Shepherd, B., and Whittington, J. (2006). Response of wolves to corridor restoration and human use management. Ecology and Society 11, 1–15.

Shille, V. M., Haggerty, M. A., Shackleton, C., and Lasley, B. L. (1990). Metabolites of estradiol in serum, bile, intestine and feces of the domestic cat. Theriogenology 34, 779–794.
| CrossRef | CAS |

Stahl, P., and Artois, M. (Eds.) (1991). ‘Status and Conservation of the Wildcat (Felis silvestris) in Europe and around the Mediterranean Rim.’ (Council of Europe: Strasbourg.)

Stahl, P., and Leger, F. (1992). Le chat sauvage (Felis silvestris, Schreber, 1777). In ‘Encyclopédie des Carnivores de France’. (Ed. M. Artois and H. Maurin.) pp. 1–50. (Société Française pour l’Etude et la Protection des Mammifères (S.F.E.P.M.): Bohallard, Puceul.)

Sunquist, M., and Sunquist, F. (Eds) (2002). ‘Wild Cats of the World.’ (The University of Chicago Press: Chicago, IL.)

Terio, K. A., Marker, L., and Munson, L. (2004). Evidence for chronic stress in captive but not free-ranging cheetahs (Acinonyx jubatus) based on adrenal morphology and function. Journal of Wildlife Diseases 40, 259–266.
| CAS |

Theuerkauf, J., Gula, R., Pirga, B., Tsunoda, H., Eggermann, J., Brzezowska, B., Rouys, S., and Radler, S. (2007). Human impact on wolf activity in the Bieszczady Mountains, SE Poland. Annales Zoologici Fennici 44, 225–231.

Thiel, D., Jenni-Eiermann, S., Braunisch, V., Palme, R., and Jenni, L. (2008). Ski tourism affects habitat use and evokes a physiological stress response in capercaillie Tetrao urogallus: a new methodological approach. Journal of Applied Ecology 45, 845–853.
| CrossRef |

Thomas, C. D., Cameron, A., and Green, R. E. (2004). Extinction risk from climate change. Nature 427, 145–148.
| CrossRef | CAS |

Tilman, D., Cassman, K. G., Matson, P. A., Naylor, R., and Polasky, S. (2002). Agricultural sustainability and intensive production practices. Nature 418, 671–677.
| CrossRef | CAS |

Touma, C., and Palme, R. (2005). Measuring fecal glucocorticoid metabolites in mammals and birds: the importance of validation. Annals of the New York Academy of Sciences 1046, 54–74.
| CrossRef | CAS |

Van Dyke, F. G., Broche, R. H., Shaw, H. G., Ackerman, B. B., Hemker, T. P., and Lindzey, F. G. (1986). Reactions on mountain lions to logging and human activity. The Journal of Wildlife Management 50, 95–102.
| CrossRef |

Van Meter, P. E., French, J. A., Dloniak, S. M., Watts, H. E., Kolowski, J. M., and Holekamp, K. E. (2009). Fecal glucocorticoids reflect socio-ecological and anthropogenic stressors in the lives of wild spotted hyenas. Hormones and Behavior 55, 329–337.
| CrossRef | CAS |

Verhage, H. G., Beamer, N. B., and Brenner, R. M. (1976). Plasma levels of estradiol and progesterone in the cat during polyestrus, pregnancy and pseudopregnancy. Biology of Reproduction 14, 579–585.
| CrossRef | CAS |

von der Ohe, C. G., Wasser, S. K., Hunt, K. E., and Servheen, C. (2004). Factors associated with fecal glucocorticoids in Alaskan brown bears (Ursus arctos horribilis). Physiological and Biochemical Zoology 77, 313–320.
| CrossRef | CAS |

Wielebnowski, N. C., Fletchall, N., Carlstead, K., Busso, J. M., and Brown, J. L. (2002). Noninvasive assessment of adrenal activity associated with husbandry and behavioral factors in the North American clouded leopard population. Zoo Biology 21, 77–98.
| CrossRef |

Young, K. M., Walker, S. L., Lanthier, C., Waddell, W. T., Monfort, S. L., and Brown, J. L. (2004). Noninvasive monitoring of adrenocortical activity in carnivores by faecal glucocorticoid analyses. General and Comparative Endocrinology 137, 148–165.
| CrossRef | CAS |