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RESEARCH ARTICLE

Dual-purpose cereals: can the relative influences of management and environment on crop recovery and grain yield be dissected?

Matthew T. Harrison A B C D , John R. Evans B , Hugh Dove A and Andrew D. Moore A
+ Author Affiliations
- Author Affiliations

A CSIRO Plant Industry, GPO Box 1600, Canberra, ACT 2601, Australia.

B The Australian National University, Research School of Biology, Canberra, ACT 0200, Australia.

C Present address: INRA, UMR 0759, Laboratoire d’Ecophysiologie des Plantes sous Stress Environnementaux, IBIP – Bât. 7, 2 Place Viala, F-34060 Montpellier, France.

D Corresponding author. Email: matthew.harrison@supagro.inra.fr

Crop and Pasture Science 62(11) 930-946 https://doi.org/10.1071/CP11066
Submitted: 14 March 2011  Accepted: 17 November 2011   Published: 21 December 2011

Abstract

Growing cereal crops for the dual-purposes (DP) of livestock forage during the early vegetative stages and harvesting grain at maturity has been practised for decades. It follows that scientific experiments using DP crops are nearly as old. A survey of more than 270 DP crop experiments revealed that the average effect of crop defoliation on grain yield (GY) was −7 ± 25% (range –35 to 75%). In light of these results, the first purpose of this review was to assess how alternative crop and grazing management regimes affected forage production and GY. Management techniques in order of decreasing importance likely to maximise grain production include (i) terminating grazing at or before GS 30, (ii) matching crop phenology to environment type, (iii) sowing DP crops 2–4 weeks earlier than corresponding sowing dates of grain-only crops, and (iv) ensuring good crop establishment before commencement of grazing. The second aim was to identify the environmental and biotic mechanisms underpinning crop responses to grazing, and to identify crop traits that would be most conducive to minimising yield penalty. A variety of mechanisms increased GY after grazing. Under favourable conditions, increased GY of grazed crops occurred via reduced lodging, mitigation of foliar disease and rapid leaf area recovery after grazing. Under stressful conditions, increased yields of grazed crops were caused by reduced transpiration and conservation of soil water, delayed phenology (frost avoidance at anthesis), and high ability to retranslocate stem reserves to grain. Yield reductions caused by grazing were associated with (i) frost damage soon after grazing, (ii) poor leaf area development or (iii) delayed maturation, which led to water or temperature stress around anthesis, culminating in increased rates of green area senescence and decreased duration of grain-filling. The third aim was to examine the role of simulation models in dissecting the effects of environment from management on crop physiology. Simulation studies of DP crops have extended the results from experimental studies, confirming that forage production increases with earlier sowing, but have also revealed that chances of liveweight gain increase with earlier sowing. Recent modelling demonstrates that potential for inclusion of DP crops into traditional grain-only systems is high, except where growing-season rainfall is <300 mm. Prospective research involving crop defoliation should focus on crop recovery, specifically (i) the effects of defoliation on phenology, (ii) the time-course of leaf area recovery and dry matter partitioning, and/or (iii) development of crop-grazing models, for these three areas will be most conducive to increasing the understanding of crop responses to grazing, thereby leading to better management guidelines.

Additional keywords: defoliation, herbivory, leaf area, radiation, soil water, transpiration.


References

Amjad M, Sharma DL, Curtis B, Anderson WK (2006) Dual-purpose long season winter wheats to improve productivity in Western Australia. In ‘Proceedings of the 13th Australian Society of Agronomy Conference’. September 2006, Perth, W. Aust. (Eds N Turner, T Acuna) (Australian Society of Agronomy: Perth)

Anderson WK (1985) Production of green feed and grain from grazed barley in northern Syria. Field Crops Research 10, 57–75.
Production of green feed and grain from grazed barley in northern Syria.Crossref | GoogleScholarGoogle Scholar |

Arzadun MJ, Arroquy JI, Laborde HE, Brevedan RE (2003) Grazing pressure on beef and grain production of dual-purpose wheat in Argentina. Agronomy Journal 95, 1157–1162.
Grazing pressure on beef and grain production of dual-purpose wheat in Argentina.Crossref | GoogleScholarGoogle Scholar |

Asghar M, Ingram BF (1993) Effects of defoliation on dryland wheat production in central Queensland. Australian Journal of Experimental Agriculture 33, 349–351.
Effects of defoliation on dryland wheat production in central Queensland.Crossref | GoogleScholarGoogle Scholar |

Baumhardt RL, Schwartz RC, Greene LW, MacDonald JC (2009) Cattle gain and crop yield for a dryland wheat–sorghum–fallow rotation. Agronomy Journal 101, 150–158.
Cattle gain and crop yield for a dryland wheat–sorghum–fallow rotation.Crossref | GoogleScholarGoogle Scholar |

Benjamin RW, Chen M, Seligman NG, Wallach D, Hadad MJA (1978) Primary production of grazed annual natural pasture and of grazed wheat in a semi-arid region of Israel. Agricultural Systems 3, 205–220.
Primary production of grazed annual natural pasture and of grazed wheat in a semi-arid region of Israel.Crossref | GoogleScholarGoogle Scholar |

Bonachela S, Orgaz F, Fereres E (1995a) Winter cereals grown for grain and for the dual purpose of forage plus grain. 1. Production. Field Crops Research 44, 1–11.
Winter cereals grown for grain and for the dual purpose of forage plus grain. 1. Production.Crossref | GoogleScholarGoogle Scholar |

Bonachela S, Orgaz F, Fereres E (1995b) Winter cereals grown for grain and for the dual purpose of forage plus grain. 2. Water use and water-use efficiency. Field Crops Research 44, 13–24.
Winter cereals grown for grain and for the dual purpose of forage plus grain. 2. Water use and water-use efficiency.Crossref | GoogleScholarGoogle Scholar |

Borrell AK, Hammer GL (2000) Nitrogen dynamics and the physiological basis of stay-green in sorghum. Crop Science 40, 1295–1307.
Nitrogen dynamics and the physiological basis of stay-green in sorghum.Crossref | GoogleScholarGoogle Scholar |

Branson FA (1953) Two new factors affecting resistance of grasses to grazing. Journal of Range Management 6, 165–171.
Two new factors affecting resistance of grasses to grazing.Crossref | GoogleScholarGoogle Scholar |

Christiansen S, Svejcar T, Phillips WA (1989) Spring and fall cattle grazing effects on components and total grain yield of winter wheat. Agronomy Journal 81, 145–150.
Spring and fall cattle grazing effects on components and total grain yield of winter wheat.Crossref | GoogleScholarGoogle Scholar |

Condon AG, Richards RA, Farquhar GD (1993) Relationships between carbon-isotope discrimination, water-use efficiency and transpiration efficiency for dryland wheat. Australian Journal of Agricultural Research 44, 1693–1711.
Relationships between carbon-isotope discrimination, water-use efficiency and transpiration efficiency for dryland wheat.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DyaK2cXhtVais78%3D&md5=b91bc7ffd8bf497a12ca060b40acf296CAS |

Cooper PJM, Gregory PJ, Tully D, Harris HC (1987) Improving water-use efficiency of annual crops in the rain-fed farming systems of West Asia and North-Africa. Experimental Agriculture 23, 113–158.
Improving water-use efficiency of annual crops in the rain-fed farming systems of West Asia and North-Africa.Crossref | GoogleScholarGoogle Scholar |

Cutler GH, Pavez DS, Mulvey RR (1949) The effect of clipping to simulate pasturing winter wheat on the growth, yield and quality of the crop. Agronomy Journal 41, 169–173.
The effect of clipping to simulate pasturing winter wheat on the growth, yield and quality of the crop.Crossref | GoogleScholarGoogle Scholar |

Dann PR (1968) Effect of clipping on yield of wheat. Australian Journal of Experimental Agriculture 8, 731–735.
Effect of clipping on yield of wheat.Crossref | GoogleScholarGoogle Scholar |

Dann PR, Axelsen A, Dear BS, Williams ER, Edwards CBH (1983) Herbage, grain and animal production from winter grazed cereal crops. Australian Journal of Experimental Agriculture and Animal Husbandry 23, 154–161.
Herbage, grain and animal production from winter grazed cereal crops.Crossref | GoogleScholarGoogle Scholar |

Dann PR, Axelsen A, Edwards CBH (1977) The grain yield of winter grazed crops. Australian Journal of Experimental Agriculture 17, 452–461.
The grain yield of winter grazed crops.Crossref | GoogleScholarGoogle Scholar |

Davidson JL, Jones DB, Christian KR (1990) Winter feed production and grain yield in mixtures of spring and winter wheats. Australian Journal of Agricultural Research 41, 1–18.
Winter feed production and grain yield in mixtures of spring and winter wheats.Crossref | GoogleScholarGoogle Scholar |

de Mello ACL, Pedreira CGS (2004) Morphological responses of irrigated Tanzaniagrass (Panicum maximum jacq. cv. Tanzania-1) to grazing intensity under rotational stocking. Revista Brasileira De Zootecnia – Brazilian Journal of Animal Science 33, 282–289.

Dean G (2007) Optimal plant density and the effect of grazing on grain yield of early sown wheat – Epping Forest, Tasmania. Southern Farming Systems. Available at: www.ccmaknowledgebase.vic.gov.au/resources/16.pdf (accessed 3 December 2011).

Dean G, Davey B, Munford S (2006) Effect of fodder cuts on grain yield of early sown wheat and triticale – Perth, Tas. Southern Farming Systems. Available at: www.sfs.org.au/G&G/2006TrialResults/GG6.htm (accessed 12 November 2007).

Doole GJ, Bathgate AD, Robertson MJ (2009) Economic value of grazing vegetative wheat (Triticum aestivum L.) crops in mixed-farming systems of Western Australia. Animal Production Science 49, 807–815.
Economic value of grazing vegetative wheat (Triticum aestivum L.) crops in mixed-farming systems of Western Australia.Crossref | GoogleScholarGoogle Scholar |

Dove H, Holst PJ, Stanley DF, Flint PW (2002) Grazing value of dual-purpose wheats for young sheep. Animal Production in Australia 24, 53–56.

Dove H, McMullen KG (2009) Diet selection, herbage intake and liveweight gain in young sheep grazing dual-purpose wheats and sheep responses to mineral supplements. Animal Production Science 49, 749–758.
Diet selection, herbage intake and liveweight gain in young sheep grazing dual-purpose wheats and sheep responses to mineral supplements.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DC%2BD1MXhtFelsbvL&md5=c4f87b5f5e5178032e22509d88d12ea6CAS |

Duggan BL, Richards RA, van Herwaarden AF (2005) Agronomic evaluation of a tiller inhibition gene (tin) in wheat. II. Growth and partitioning of assimilate. Australian Journal of Agricultural Research 56, 179–186.
Agronomic evaluation of a tiller inhibition gene (tin) in wheat. II. Growth and partitioning of assimilate.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DC%2BD2MXhvFyku7o%3D&md5=8ed2cab6db038f846498fe48c1194538CAS |

Duncan WG (1971) Leaf angles, leaf area, and canopy photosynthesis. Crop Science 11, 482–485.
Leaf angles, leaf area, and canopy photosynthesis.Crossref | GoogleScholarGoogle Scholar |

Dunin F, Passioura J (2006) Prologue: amending agricultural water use to maintain production while affording environmental protection through control of outflow. Australian Journal of Agricultural Research 57, 251–255.
Prologue: amending agricultural water use to maintain production while affording environmental protection through control of outflow.Crossref | GoogleScholarGoogle Scholar |

Dunphy DJ, Holt EC, McDaniel ME (1984) Leaf area and dry matter accumulation of wheat following forage removal. Agronomy Journal 76, 871–874.
Leaf area and dry matter accumulation of wheat following forage removal.Crossref | GoogleScholarGoogle Scholar |

Dunphy DJ, McDaniel ME, Holt EC (1982) Effect of forage utilization on wheat grain yield. Crop Science 22, 106–109.
Effect of forage utilization on wheat grain yield.Crossref | GoogleScholarGoogle Scholar |

Edwards JT, Carver BF, Horn GW, Payton ME (2011) Impact of dual-purpose management on wheat grain yield. Crop Science 47, 2074–2077.

Ehdaie B, Alloush GA, Madore MA, Waines JG (2006) Genotypic variation for stem reserves and mobilization in wheat: I. Postanthesis changes in internode dry matter. Crop Science 46, 735–746.
Genotypic variation for stem reserves and mobilization in wheat: I. Postanthesis changes in internode dry matter.Crossref | GoogleScholarGoogle Scholar |

Epplin FM, Hossain I, Krenzer EG (2000) Winter wheat fall-winter forage yield and grain yield response to planting date in a dual-purpose system. Agricultural Systems 63, 161–173.
Winter wheat fall-winter forage yield and grain yield response to planting date in a dual-purpose system.Crossref | GoogleScholarGoogle Scholar |

Evans LT (1960) Inflorescence initiation in Lolium temulentum L. 1. Effect of plant age and leaf area on sensitivity to photo-periodic induction. Australian Journal of Biological Sciences 13, 123–131.

Evans LT, Wardlaw IF, Fischer RA (1975) Wheat. In ‘Crop physiology: some case histories’. (Ed. LT Evans) pp. 101–150. (Cambridge University Press: London)

Fahnestock JT, Detling JK (2000) Morphological and physiological responses of perennial grasses to long-term grazing in the Pryor Mountains, Montana. American Midland Naturalist 143, 312–320.
Morphological and physiological responses of perennial grasses to long-term grazing in the Pryor Mountains, Montana.Crossref | GoogleScholarGoogle Scholar |

Fahnestock JT, Knapp AK (1994) Plant responses to selective grazing by bison – interactions between light, herbivory and water-stress. Vegetatio 115, 123–131.

Fieser BG, Horn GW, Edwards JT, Krezner EG (2006) Timing of grazing termination in dual-purpose winter wheat enterprises. The Professional Animal Scientist 22, 210–216.

Fischer RA (1979) Growth and water limitation to dryland wheat yield in Australia – a physiological framework. Journal of the Australian Institute of Agricultural Science 45, 83–94.

Fischer RA (2007) Understanding the physiological basis of yield potential in wheat. The Journal of Agricultural Science 145, 99–113.
Understanding the physiological basis of yield potential in wheat.Crossref | GoogleScholarGoogle Scholar |

Francia E, Pecchioni N, Nicosia OLD, Paoletta G, Taibi L, Franco V, Odoardi M, Stanca AM, Delogu G (2006) Dual-purpose barley and oat in a Mediterranean environment. Field Crops Research 99, 158–166.
Dual-purpose barley and oat in a Mediterranean environment.Crossref | GoogleScholarGoogle Scholar |

French RJ, Schultz JE (1984) Water-use efficiency of wheat in a Mediterranean-type environment. 1. The relation between yield, water-use and climate. Australian Journal of Agricultural Research 35, 743–764.
Water-use efficiency of wheat in a Mediterranean-type environment. 1. The relation between yield, water-use and climate.Crossref | GoogleScholarGoogle Scholar |

García del Moral LF (1992) Leaf area, grain yield and yield components following forage removal in triticale. Journal of Agronomy and Crop Science - Zeitschrift Fur Acker Und Pflanzenbau 168, 100–107.
Leaf area, grain yield and yield components following forage removal in triticale.Crossref | GoogleScholarGoogle Scholar |

Gastal F, Lemaire G (2002) N uptake and distribution in crops: an agronomical and ecophysiological perspective. Journal of Experimental Botany 53, 789–799.
N uptake and distribution in crops: an agronomical and ecophysiological perspective.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DC%2BD38XivFSntL0%3D&md5=6e66d093e32d62779009b514ca7de02dCAS |

Gautier H, Varlet-Grancher C, Hazard L (1999) Tillering responses to the light environment and to defoliation in populations of perennial ryegrass (Lolium perenne L.) selected for contrasting leaf length. Annals of Botany 83, 423–429.
Tillering responses to the light environment and to defoliation in populations of perennial ryegrass (Lolium perenne L.) selected for contrasting leaf length.Crossref | GoogleScholarGoogle Scholar |

Hacking C (2006) Dual purpose cereal variety evaluation – Bairnsdale, Vic. Available at: www.ccmaknowledgebase.vic.gov.au/resources/9.pdf (accessed 14 November 2007).

Hammer GL, Wright GC (1994) A theoretical analysis of nitrogen and radiation effects on radiation use efficiency in peanut. Australian Journal of Agricultural Research 45, 575–589.
A theoretical analysis of nitrogen and radiation effects on radiation use efficiency in peanut.Crossref | GoogleScholarGoogle Scholar |

Harrison MT, Kelman WM, Moore AD, Evans JR (2010) Grazing winter wheat relieves plant water stress and transiently enhances photosynthesis. Functional Plant Biology 37, 726–736.
Grazing winter wheat relieves plant water stress and transiently enhances photosynthesis.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DC%2BC3cXpt1Ckt7Y%3D&md5=1bde609c08d599d7fc931960ea20d3b2CAS |

Harrison MT, Evans JR, Dove H, Moore AD (2011a) Recovery dynamics of rainfed winter wheat after livestock grazing. 1. Growth rates, grain yields, soil water use and water-use efficiency. Crop & Pasture Science 62, 947–959.

Harrison MT, Evans JR, Dove H, Moore AD (2011b) Recovery dynamics of rainfed winter wheat after livestock grazing. 2. Light interception, radiation-use efficiency and dry-matter partitioning. Crop & Pasture Science 62, 960–971.

Hatfield JL, Sauer TJ, Prueger JH (2001) Managing soils to achieve greater water use efficiency: a review. Agronomy Journal 93, 271–280.
Managing soils to achieve greater water use efficiency: a review.Crossref | GoogleScholarGoogle Scholar |

Hofstede RGM, Chilito EJ, Sandovals EM (1995) Vegetative structure, microclimate, and leaf growth of a paramo tussock grass species, in undisturbed, burned and grazed conditions. Vegetatio 119, 53–65.

Holman JD, Thompson CR, Hale RL, Schlegel AJ (2009) Grazing effects on yield and quality of hard red and hard white winter wheat. Agronomy Journal 101, 775–788.
Grazing effects on yield and quality of hard red and hard white winter wheat.Crossref | GoogleScholarGoogle Scholar |

Hubbard VC, Harper HJ (1949) Effect of clipping small grains on composition and yield of forage and grain. Agronomy Journal 41, 85–92.
Effect of clipping small grains on composition and yield of forage and grain.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DyaH1MXivFOguw%3D%3D&md5=3f2cde399b7cbab23e3277ee7f6ab4fbCAS |

Kelman WM, Dove H (2007) Effects of a spring-sown brassica crop on lamb performance and on subsequent establishment and grain yield of dual-purpose winter wheat and oat crops. Australian Journal of Experimental Agriculture 47, 815–824.
Effects of a spring-sown brassica crop on lamb performance and on subsequent establishment and grain yield of dual-purpose winter wheat and oat crops.Crossref | GoogleScholarGoogle Scholar |

Kelman WM, Dove H (2009) Growth and phenology of winter wheat and oats in a dual-purpose management system. Crop & Pasture Science 60, 921–932.
Growth and phenology of winter wheat and oats in a dual-purpose management system.Crossref | GoogleScholarGoogle Scholar |

Kelman WM, Harrison MT (2010) Interactions between plant density and grazing in cereals under dual-purpose management. In ‘Food Security from Sustainable Agriculture. Proceedings of the 15th Australian Society of Agronomy Conference’. 15–19 November 2010, Lincoln University, New Zealand. (Eds H Dove, RA Culvenor) (Australian Society of Agronomy: Lincoln, New Zealand)

Kemanian AR, Stockle CO, Huggins DR (2005) Transpiration-use efficiency of barley. Agricultural and Forest Meteorology 130, 1–11.

Khalil IH, Carver BF, Krenzer EG, MacKown CT, Horn GW (2002) Genetic trends in winter wheat yield and test weight under dual-purpose and grain-only management systems. Crop Science 42, 710–715.
Genetic trends in winter wheat yield and test weight under dual-purpose and grain-only management systems.Crossref | GoogleScholarGoogle Scholar |

Kirkegaard JA, Lilley JM, Howe GN, Graham JM (2007) Impact of subsoil water use on wheat yield. Australian Journal of Agricultural Research 58, 303–315.
Impact of subsoil water use on wheat yield.Crossref | GoogleScholarGoogle Scholar |

Large EC (1954) Growth stages in cereals – illustration of the Feekes scale. Plant Pathology 3, 128–129.
Growth stages in cereals – illustration of the Feekes scale.Crossref | GoogleScholarGoogle Scholar |

Lovett JV, Matheson EM (1974) Cereals for winter grazing on the northern tablelands of New South Wales. Australian Journal of Experimental Agriculture and Animal Husbandry 14, 790–795.
Cereals for winter grazing on the northern tablelands of New South Wales.Crossref | GoogleScholarGoogle Scholar |

Lyon DJ, Baltensperger DD, Siles M (2001) Wheat grain and forage yields are affected by planting and harvest dates in the Central Great Plains. Crop Science 41, 488–492.
Wheat grain and forage yields are affected by planting and harvest dates in the Central Great Plains.Crossref | GoogleScholarGoogle Scholar |

MacKown CT, Carver BF (2005) Fall forage biomass and nitrogen composition of winter wheat populations selected from grain-only and dual-purpose environments. Crop Science 45, 322–328.
Fall forage biomass and nitrogen composition of winter wheat populations selected from grain-only and dual-purpose environments.Crossref | GoogleScholarGoogle Scholar |

MacKown CT, Carver BF (2007) Nitrogen use and biomass distribution in culms of winter wheat populations selected from grain-only and dual-purpose systems. Crop Science 47, 350–358.
Nitrogen use and biomass distribution in culms of winter wheat populations selected from grain-only and dual-purpose systems.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DC%2BD2sXjsF2msro%3D&md5=bbd9064f52e7b9cbb4b639d9946b3276CAS |

Matches AG (1992) Plant-response to grazing – a review. Journal of Production Agriculture 5, 1–7.

Mauget S, Zhang J, Ko J (2009) The value of ENSO forecast information to dual-purpose winter wheat production in the US southern high plains. Journal of Applied Meteorology and Climatology 48, 2100–2117.
The value of ENSO forecast information to dual-purpose winter wheat production in the US southern high plains.Crossref | GoogleScholarGoogle Scholar |

McMullen KG, Virgona JM (2009) Dry matter production and grain yield from grazed wheat in southern New South Wales. Animal Production Science 49, 769–776.
Dry matter production and grain yield from grazed wheat in southern New South Wales.Crossref | GoogleScholarGoogle Scholar |

Messina C, Hammer G, Zhanshan D, Podlich D, Cooper M (2009) Modelling crop improvement in a G × E × M framework via gene–trait–phenotype relationships. In ‘Crop physiology: interfacing with genetic improvement and agronomy’. (Eds VO Sadras, D Calderini) pp. 235–266. (Elsevier Inc.: The Netherlands)

Miller DR, Dean GJ, Ball PD (2010) Influence of end-grazing forage residual and grazing management on lamb growth performance and crop yield from irrigated dual-purpose winter wheat. Animal Production Science 50, 508–512.
Influence of end-grazing forage residual and grazing management on lamb growth performance and crop yield from irrigated dual-purpose winter wheat.Crossref | GoogleScholarGoogle Scholar |

Miller GL, Joost RE, Harrison SA (1993) Forage and grain yields of wheat and triticale as affected by forage management practices. Crop Science 33, 1070–1075.
Forage and grain yields of wheat and triticale as affected by forage management practices.Crossref | GoogleScholarGoogle Scholar |

Milroy SP, Goyne PJ (1995) Leaf area development in barley – model construction and response to soil moisture status. Australian Journal of Agricultural Research 46, 845–860.

Miralles DJ, Slafer GA (1997) Radiation interception and radiation use efficiency of near-isogenic wheat lines with different height. Euphytica 97, 201–208.
Radiation interception and radiation use efficiency of near-isogenic wheat lines with different height.Crossref | GoogleScholarGoogle Scholar |

Moore AD (2009) Opportunities and trade-offs in dual-purpose cereals across the southern Australian mixed-farming zone: a modelling study. Animal Production Science 49, 759–768.
Opportunities and trade-offs in dual-purpose cereals across the southern Australian mixed-farming zone: a modelling study.Crossref | GoogleScholarGoogle Scholar |

Moore AD, Bell LW, Revell DK (2009) Feed-gaps in mixed-farming systems: insights from the Grain & Graze program. Animal Production Science 49, 736–748.
Feed-gaps in mixed-farming systems: insights from the Grain & Graze program.Crossref | GoogleScholarGoogle Scholar |

Muir CE, Virgona JM, Angus JF (2007) Grazing effects on the retranslocation of assimilates during the grain filling of wheat. Available at: www.regional.org.au/au/asa/2006/poster/pests/4672_muirc.htm (accessed 18 November 2007).

Nicholson C (2006) Effect of grazing on the grain yield and quality of seven cereals – Inverleigh, Vic. In ‘Proceedings of the 13th Australian Agronomy Conference’. Perth, Western Australia. (Eds NC Turner, T Acuna, RC Johnson) pp. 184–186. (Southern Farming Systems, Australia: Newtown)

Nowak RS, Caldwell MM (1984) A test of compensatory photosynthesis in the field – implications for herbivory tolerance. Oecologia 61, 311–318.
A test of compensatory photosynthesis in the field – implications for herbivory tolerance.Crossref | GoogleScholarGoogle Scholar |

Noy-Meir I, Briske DD (2002) Response of wild wheat populations to grazing in Mediterranean grasslands: the relative influence of defoliation, competition, mulch and genotype. Journal of Applied Ecology 39, 259–278.
Response of wild wheat populations to grazing in Mediterranean grasslands: the relative influence of defoliation, competition, mulch and genotype.Crossref | GoogleScholarGoogle Scholar |

Nuske K, Hunt J, Best F (2009) Grazing cereals. In ‘BCG 2009 season research results’. pp. 46–51. (Birchip Cropping Group: Melbourne)

Passioura J (2006) Increasing crop productivity when water is scarce – from breeding to field management. Agricultural Water Management 80, 176–196.
Increasing crop productivity when water is scarce – from breeding to field management.Crossref | GoogleScholarGoogle Scholar |

Pumphrey FV (1970) Semidwarf winter wheat response to early spring clipping and grazing. Agronomy Journal 62, 641–643.
Semidwarf winter wheat response to early spring clipping and grazing.Crossref | GoogleScholarGoogle Scholar |

Ramos JM, Delmoral MBG, Marinetto J, Delmoral LFG (1993) Sowing date and cutting frequency effects on triticale forage and grain production. Crop Science 33, 1312–1315.
Sowing date and cutting frequency effects on triticale forage and grain production.Crossref | GoogleScholarGoogle Scholar |

Ramos JM, García del Moral LF, Boujenna A, Serra J, Insa JA, Royo C (1996) Grain yield, biomass and leaf area of triticale in response to sowing date and cutting stage in three contrasting Mediterranean environments. The Journal of Agricultural Science 126, 253–258.
Grain yield, biomass and leaf area of triticale in response to sowing date and cutting stage in three contrasting Mediterranean environments.Crossref | GoogleScholarGoogle Scholar |

Redmon LA, Krenzer EG, Bernardo DJ, Horn GW (1996) Effect of wheat morphological stage at grazing termination on economic return. Agronomy Journal 88, 94–97.
Effect of wheat morphological stage at grazing termination on economic return.Crossref | GoogleScholarGoogle Scholar |

Rice EL, Parenti RL (1978) Causes of decreases in productivity in undisturbed tall grass prairie. American Journal of Botany 65, 1091–1097.
Causes of decreases in productivity in undisturbed tall grass prairie.Crossref | GoogleScholarGoogle Scholar |

Richards RA (2000) Selectable traits to increase crop photosynthesis and yield of grain crops. Journal of Experimental Botany 51, 447–458.
Selectable traits to increase crop photosynthesis and yield of grain crops.Crossref | GoogleScholarGoogle Scholar | 1:CAS:528:DC%2BD3cXhsVOjsrw%3D&md5=195fb7696213e4e872f32d2f307599f9CAS |

Richards RA, Rebetzke GJ, Condon AG, van Herwaarden AF (2002) Breeding opportunities for increasing the efficiency of water use and crop yield in temperate cereals. Crop Science 42, 111–121.
Breeding opportunities for increasing the efficiency of water use and crop yield in temperate cereals.Crossref | GoogleScholarGoogle Scholar |

Rotundo JL, Aguiar MR (2008) Herbivory resistance traits in populations of Poa ligularis subjected to historically different sheep grazing pressure in Patagonia. Plant Ecology 194, 121–133.
Herbivory resistance traits in populations of Poa ligularis subjected to historically different sheep grazing pressure in Patagonia.Crossref | GoogleScholarGoogle Scholar |

Royo C (1997) Grain yield and yield components as affected by forage removal in winter and spring triticale. Grass and Forage Science 52, 63–72.
Grain yield and yield components as affected by forage removal in winter and spring triticale.Crossref | GoogleScholarGoogle Scholar |

Royo C (1999) Plant recovery and grain yield formation in barley and triticale following forage removal at two cutting stages. Journal of Agronomy & Crop Science 182, 175–184.
Plant recovery and grain yield formation in barley and triticale following forage removal at two cutting stages.Crossref | GoogleScholarGoogle Scholar |

Royo C, Pares D (1996) Yield and quality of winter and spring triticales for forage and grain. Grass and Forage Science 51, 449–455.
Yield and quality of winter and spring triticales for forage and grain.Crossref | GoogleScholarGoogle Scholar |

Royo C, Insa JA, Boujenna A, Ramos JM, Montesinos E, García del Moral LF (1994) Yield and quality of spring triticale used for forage and grain as influenced by sowing date and cutting stage. Field Crops Research 37, 161–168.
Yield and quality of spring triticale used for forage and grain as influenced by sowing date and cutting stage.Crossref | GoogleScholarGoogle Scholar |

Royo C, Lopez A, Serra J, Tribo F (1997) Effect of sowing date and cutting stage on yield and quality of irrigated barley and triticale used for forage and grain. Journal of Agronomy & Crop Science 179, 227–234.
Effect of sowing date and cutting stage on yield and quality of irrigated barley and triticale used for forage and grain.Crossref | GoogleScholarGoogle Scholar |

Royo C, Romagosa I (1996) Effect of forage removal at the first detectable node stage on the growth of winter and spring triticale. Grass and Forage Science 51, 170–179.
Effect of forage removal at the first detectable node stage on the growth of winter and spring triticale.Crossref | GoogleScholarGoogle Scholar |

Royo C, Tribo F (1997a) Triticale and barley for grain and for dual-purpose (forage plus grain) in a Mediterranean-type environment. 1. Growth analyses. Australian Journal of Agricultural Research 48, 411–421.
Triticale and barley for grain and for dual-purpose (forage plus grain) in a Mediterranean-type environment. 1. Growth analyses.Crossref | GoogleScholarGoogle Scholar |

Royo C, Tribo F (1997b) Triticale and barley for grain and for dual-purpose (forage plus grain) in a Mediterranean-type environment. 2. Yield, yield components, and quality. Australian Journal of Agricultural Research 48, 423–432.
Triticale and barley for grain and for dual-purpose (forage plus grain) in a Mediterranean-type environment. 2. Yield, yield components, and quality.Crossref | GoogleScholarGoogle Scholar |

Scott WR, Hines SE (1991) Effects of grazing on grain yield of winter barley and triticale – the position of the apical dome relative to the soil surface. New Zealand Journal of Agricultural Research 34, 177–184.

Sharrow SH, Motazedian I (1987) Spring grazing effects on components of winter-wheat yield. Agronomy Journal 79, 502–504.
Spring grazing effects on components of winter-wheat yield.Crossref | GoogleScholarGoogle Scholar |

Simpson RJ, Culvenor RA (1987) Photosynthesis, carbon partitioning and herbage yield. In ‘Temperate pastures – their production, use and management’. (Eds JL Wheeler, CJ Pearson, GE Robards) pp. 103–118. (Australian Wool Corporation Technical Publication: Melbourne)

Slafer GA, Rawson HM (1994) Sensitivity of wheat phasic development to major environmental factors – a reexamination of some assumptions made by physiologists and modelers. Australian Journal of Plant Physiology 21, 393–426.
Sensitivity of wheat phasic development to major environmental factors – a reexamination of some assumptions made by physiologists and modelers.Crossref | GoogleScholarGoogle Scholar |

Sprague MA (1954) The effect of grazing management on forage and grain production from rye, wheat and oats. Agronomy Journal 46, 29–33.
The effect of grazing management on forage and grain production from rye, wheat and oats.Crossref | GoogleScholarGoogle Scholar |

Stone PJ, Sorensen IB, Jamieson PD (1999) Effect of soil temperature on phenology, canopy development, biomass and yield of maize in a cool-temperate climate. Field Crops Research 63, 169–178.
Effect of soil temperature on phenology, canopy development, biomass and yield of maize in a cool-temperate climate.Crossref | GoogleScholarGoogle Scholar |

Swanson AF (1935) Pasturing winter wheat in Kansas. Kansas Agricultural Experiment Station Bulletin 271.

Taylor KW, Epplin FM, Brorsen BW, Fieser BG, Horn GW (2010) Optimal grazing termination date for dual-purpose winter wheat production. Journal of Agricultural and Applied Economics 42, 87–103.

van Herwaarden AF (1996) Haying-off in wheat: enduring myth or current problem? In ‘Agronomy – Science with its sleeves rolled up. Proceedings of the 8th Australian Agronomy Conference’. Toowoomba, Queensland, 30 January–2 February 1996. (Eds DL Michalk, JE Pratley) pp. 566–569. (Australian Society of Agronomy: Toowoomba, Qld)

Virgona JM, Gummer FAJ, Angus JF (2006) Effects of grazing on wheat growth, yield, development, water use, and nitrogen use. Australian Journal of Agricultural Research 57, 1307–1319.
Effects of grazing on wheat growth, yield, development, water use, and nitrogen use.Crossref | GoogleScholarGoogle Scholar |

Wan CG, Sosebee RE (1998) Tillering responses to red: far-red light ratio during different phenological stages in Eragrostis curvula. Environmental and Experimental Botany 40, 247–254.
Tillering responses to red: far-red light ratio during different phenological stages in Eragrostis curvula.Crossref | GoogleScholarGoogle Scholar |

Winter SR, Musick JT (1991) Grazed wheat grain yield relationships. Agronomy Journal 83, 130–135.
Grazed wheat grain yield relationships.Crossref | GoogleScholarGoogle Scholar |

Winter SR, Thompson EK (1987) Grazing duration effects on wheat growth and grain yield. Agronomy Journal 79, 110–114.
Grazing duration effects on wheat growth and grain yield.Crossref | GoogleScholarGoogle Scholar |

Winter SR, Thompson EK (1990) Grazing winter wheat. I. Response of semidwarf cultivars to grain and grazed production systems. Agronomy Journal 82, 33–37.
Grazing winter wheat. I. Response of semidwarf cultivars to grain and grazed production systems.Crossref | GoogleScholarGoogle Scholar |

Winter SR, Thompson EK, Musick JT (1990) Grazing winter wheat. II. Height effects on response to production systems. Agronomy Journal 82, 37–41.
Grazing winter wheat. II. Height effects on response to production systems.Crossref | GoogleScholarGoogle Scholar |

Yau SK (2003) Yields of early planted barley after clipping or grazing in a semiarid area. Agronomy Journal 95, 821–827.
Yields of early planted barley after clipping or grazing in a semiarid area.Crossref | GoogleScholarGoogle Scholar |

Zadoks JC, Chang TT, Konzak CF (1974) Decimal code for growth stages of cereals. Weed Research 14, 415–421.
Decimal code for growth stages of cereals.Crossref | GoogleScholarGoogle Scholar |

Zhang XC (2010) Optimizing stocking rate for maximum return to a wheat-cattle enterprise using model simulation and economics. Agronomy Journal 102, 197–209.
Optimizing stocking rate for maximum return to a wheat-cattle enterprise using model simulation and economics.Crossref | GoogleScholarGoogle Scholar |

Zhu GX, Midmore DJ, Radford BJ, Yule DF (2004) Effect of timing of defoliation on wheat (Triticum aestivum) in central Queensland. 1. Crop response and yield. Field Crops Research 88, 211–226.
Effect of timing of defoliation on wheat (Triticum aestivum) in central Queensland. 1. Crop response and yield.Crossref | GoogleScholarGoogle Scholar |