#NEXUS 
[MacClade 4.08 registered to Michelle DaCosta, Wake Forest University]


BEGIN TAXA;
	DIMENSIONS  NTAX=31;
	TAXLABELS
	'Argina astrea MD65/MD151'
	'Argina leonina MD64/MD153'
	'Mangina argus MD66/MD152'
	'Pitasila abraxoides 4031/2097'
	'Pitasila leucospilota 4066/4067'
	'Pitasila selecta 4052/4068'
	Utetheisa_amhara_4275
	'Utetheisa bella 213/295'
	'Utetheisa clareae 2430/1597'
	Utetheisa_devriesi_1965
	'Utetheisa dorsifumata MD317/MD318'
	'Utetheisa elata MD219/MD272'
	'Utetheisa galapagensis MD259/MD296'
	'Utetheisa lotrix MD214/MD267'
	Utetheisa_lotrix_stigmata_1534
	'Utetheisa lactea MD315/MD316'
	'Utetheisa lactea MD368/MD400'
	'Utetheisa ornatrix MD215/MD339'
	'Utetheisa perryi MD413/MD415'
	'Utetheisa pulchella MD246/MD355'
	'Utetheisa pulchella bicolor MD315/MD314'
	'Utetheisa pulchelloides MD212/MD270'
	Utetheisa_pulchelloides_aphanis_1530
	Utetheisa_pulchelloides_marshallorum_1524
	Utetheisa_pulchelloides_vaga_1529
	'Utetheisa salomonis MD307/MD309'
	'Utetheisa semara MD311/MD312'
	Atasca_pellex_MD260
	'Haploa clymene MD19/MD20'
	'Euplagia quadripunctaria MD42/MD43'
	'Tyria jacobaeae MD178/MD179';
END;


BEGIN CHARACTERS;
	DIMENSIONS  NCHAR=105;
	FORMAT SYMBOLS= " 0 1 2 3 4 5 6" MISSING=? GAP=- ;
	CHARSTATELABELS  
		1  'H1-Male antenna' / Pectinate Monofiliform Rounded_monofiliform,
  2  'H2-Female antenna' / Pectinate Monofiliform Rounded_monofiliform,
  3  'H3-Labial Palps' / Third_segment_shorter_than_or_equal_lto_second Third_segment_longer_than_second,
  4  'L1-Prothoracic tibia' / 'Unmodified,_withour_elongated_projection_proximal_to_tarsi' Modified_with_elongated_projection_proximal_to_tarsi,
  5  'L2-Length of epiphysis' / 'Long,_equal_to_or_longer_than_2/3_length_of_tibia' 'Short,_less_than_2/3_length_of_tibia',
  6  'L3-Male prothoracic tibial hair pencil' / Absent Present,
  7  'L4-Tarsal claws' / Simple Toothed,
  8  'MFW1-Sc and R1' / Separate_for_entire_distance_to_margin 'Anastomosing_for_part_of_distance_to_margin_forming_an_areole-like_cell',
  9  'MFW2-Areole' / Absent Present,
  10  'MFW3-Discal cell' / Completely_closed Partially_open,
  11  'MFW4-Vein 3A' / Absent 'Reduced,_not_reaching_margin',
  12  'MHW1- Sc and R1' / Absent Present_to_margin,
  13  'MHW2-Rs and M1' / Absent Both_present_and_free_of_each_other Short_stalked,
  14  'MHW3-M2 and M3' / 'Both_present,_and_free_of_each_other' Stalked,
  15  'MHW4-M3 and CUA1' / Separate_for_entire_distance_to_margin Stalked,
  16  'MHW5- CuA1 and CuA2' / Arising_separately_from_discal_cell Stalked,
  17  'MHW6-Scales (androconia) in anal region' / Absent Present,
  18  'FFW1-Sc and R1' / Separate_for_entire_distance_to_margin 'Anastomosing_for_part_of_distance_to_margin_forming_an_areole-like_cell',
  19  'FFW2-Areole' / Absent Present,
  20  'FFW3-Discal cell' / Completely_closed Partially_open,
  21  'FFW4-Vein 3A' / Absent 'Reduced,_not_reaching_margin',
  22  'FHW1-Sc and R1' / Absent Present_to_margin 'Reduced,_not_reaching_margin',
  23  'FHW2-Rs and M1' / Absent 'Both_present,_and_free_of_each_other' Short_stalked,
  24  'FHW3-M2 and M3' / 'Both_present,_and_free_of_each_other' Stalked 'M2_absent,_or_both_fused_along_entire_length',
  25  'FHW4-M3 and CuA1' / Separate_for_entire_distance_to_margin Stalked,
  26  'FHW5-CuA1 and CuA2' / Arising_separately_from_discal_cell Stalked Fused,
  27  'FHW6-Discal Cell' / Completely_closed Partially_open,
  28  'FHW6-Number of frenular bristles in female' / Two Three Four Five,
  29  'M1- Cephalic edge 8th abdominal segment' / Reinforced Unmodified,
  30  'M2-Length A8/A9 dorsal intersegmental membrane' / Shorter_than_7th_tergite Greater_than_or_equal_to_7th_tergite,
  31  'M3-Length of 8th tergite relative to 7th tergite' / Less_than_or_equal_to_7th_tergite Greater_than_7th_tergite,
  32  'M4-8th tergite' / Unmodified 'With_a_pair_of_antero-lateral_apodemes',
  33  'M5-Tegumen width' / 'Narrow,_less_than_or_equal_to_1mm' 'Broad,_greater_than__1mm',
  34  'M6-Articulation of tegumen with vinculum' / 'Not_fused,_with_membrane_break' 'Fused,_without_membrane_break',
  35  'M7-Uncus' / 'Thin,_slender_along_entire_length' Bulbous_just_below_apex Slightly_bulbous_just_below_apex Dorsoventrally_flattened_and_swollen_in_middle,
  36  'M8-Ornamentation of uncus' / No_setae Setae_occuring_along_entire_length Setae_occuring_from_base_to_just_below_apex,
  37  'M9-Arcticulation of uncus with tegumen' / 'Not_fused,_with_membrane_break' 'Fused,_without_membrane_break',
  38  'M10-Manica' / Membranous Reinforced_membrane,
  39  'M11-Transtilla' / Absent Present,
  40  'M12-Transtilla' / Absent Fused_to_dorsal_annellar_area Free_of_dorsal_annellar_area,
  41  'M13-Dorsal anellus' / Membranous Sclerotized,
  42  'M14-Sclerotization of dorsal anellus' / Absent Single_sclerotized_area Two_distinct_slcerotized_areas,
  43  'M15-Ventral anellus' / Membranous Sclerotized,
  44  'M16-Juxta' / Absent Present,
  45  'M17-Shape of costa' / Elongate_and_narrow_with_rodlike_apex Broad_with_square_and_flattened_apex Broad_with_rounded_dorsal_edge 'Reduced_(small)_and_flattened' Elongate_and_flattened Elongate_and_rodlike_with_spatulate_apex,
  46  'M18-Inner costal compartment' / 'Sclerotized,_unornamented' 'Scerotized,_wrinkled' 'Sclerotized,_rugose',
  47  'M19-Texture apex of costa' / Smooth Wrinkled Rugose,
  48  'M20-Outer Surface of costa' / Sclerotized Membranous,
  49  'M21-Inner saccular compartment' / 'Sclerotized,_unornamented' Membranous 'Sclerotized,_wrinkled',
  50  'M22-Texture outer apex of sacculus' / Smooth Wrinkled,
  51  'M23-Sacculus of outer valve surface with modified corematalike structure' / Absent Present,
  52  'M24-Basal Costal Process' / Absent Present,
  53  'M25-Position of basal costal process' / Absent Held_within_valve Projects_outside_valve,
  54  'M26-Orientation of basal costal process' / Dorsal Posterior Absent,
  55  'M27-Shape of basal costal process' / 'Cylindrical_and_narrow/rodlike' Flattened_and_broad Absent,
  56  'M28-Shape apex of saccus' / 'Absent,_conjunctiva' 'Rounded,_acute_angle' 'Flattened,_obtuse_angle',
  57  'M29-Condition of saccus' / Flat Inflected_towards_uncus Inflected_towards_head,
  58  'M30-Phallus' / Straight_along_entire_length With_bulge_on_ventral_side,
  59  'M31-Phallic ornamentation proximal to vesica' / Absent Present,
  60  'M32-Location of ductus ejaculatorious simplex (des)' / 'Caecum_present,_opening_dorsal' 'No_caecum,_opening_shifted_to_anterior',
  61  'M33-Vesica' / 'Unmodified,_single_lobe' 'With_lobe_projecting_laterally/dorsolaterally_on_left_side' With_lobe_projecting_dorsally 'With_dorsal,_ventral_and_posterior_lobes' With_posterior_and_ventral_lobes,
  62  'M34-Vesica with rugose patch(es)' / Absent Present,
  63  'M35-Location of rugose patch(es) on vesica' / Absent Main_lobe 'Main_lobe_+_lobe_projecting_laterally_on_left_side' 'Main_lobe_+_lobe_projecting_dorsally' 'Main_lobe_+_lobe_projecting_posteriorly' 'Two_lobes,_one_projecting_ventrally_and_the_other_posteriorly',
  64  'M36-Vesica with cornutii' / Absent Present,
  65  'M37-Arrangement of cornutti on vesica' / Absent Individually In_patches,
  66  'M38-Number of individual cornutii present' / Absent One Two Three,
  67  'M39-Number of cornutii patches' / 'Absent/Zero' One Two Three,
  68  'M40-Location of cornutii on vesica' / Absent Lobe_projecting_laterally_on_left_side Main_lobe Lobe_projecting_dorsally 'Main_lobe_+_lobe_projecting_dorsally' Lobes_projecting_dorsally_and_ventrally,
  69  'M41-Shape of cornutii' / Absesnt Spikelike Triangular Serrate,
  70  'M42-Sclerotized patches on vesica' / Absent Present,
  71  'M43-Location of sclerotized patches on vesica' / Absent On__maini_lobe_extending_beneath_dorsal_lobe 'Left_and_ventral_sides_of_main_lobe,_extending_beneath_dorsolateral_lobe',
  72  'M44-Triangular sclerotized patch on vesica' / Absent Present,
  73  'F1-Condition of ventral intersegmental membrane between A7&A8' / Unmodified With_indentations,
  74  'F2-Location of indentations in A7/A8 intersegmental membrane' / Absent Lateral_to_ostium_bursa,
  75  'F3-Condition of intersegmental membrane between A6&A7' / Unmodified With_indentations,
  76  'F4-Location of indentations in A7/A6 intersegmental membrane' / Absent 'Lateral_(closer_to_pleural_region)',
  77  'F5-Condition of 7th sternite' / Unmodified 'Modified_into_"plate"',
  78  'F6-Condition of 8th sternite' / Unmondified 'Completely_divided_mid-ventrally' 'Partially_divided_mid-ventrally' 'Associated_with_ostium_bursa_(no_clear_memb._break_bet._8S_&_OB,_instead_there_is_a_continuous_area_of_sclerotization_between_8S_and_OB)',
  79  'F7-Pseudopapillae anales' / Absent Present,
  80  'F8-Papillae anales' / Unmodified With_dorsal_extension,
  81  'F9-Posterior apophyses' / 'Short,_=_or_shorter_than_8th_sternite' 'Long,_longer_than_8th_sternite',
  82  'F10-Posterior apophyses located' / Laterally Dorsally,
  83  'F11-Anterior apophyses' / 'Short,_=_or_shorter_than_8th_sternite' 'Long,_longer_than_8th_sternite',
  84  'F12-Anterior apophyses located' / Laterally Dorsally,
  85  'F13-Angle at which 8th tergite and 8th sternite fuse' / Obtuse Acute Not_fused,
  86  'F14-Ostium bursa with sinus vaginalis' / Absent Present,
  87  'F15-ostium bursa with posterior process' / Absent Present,
  88  'F16-Post vaginal plate' / Absent Present,
  89  'F17-Dorsal intersegmental membrane A8/A9' / Unmodified With_pouch,
  90  'F18-Dorsal pheromone gland opening' / 'Not_visible,_glandular_area' Visible_as_a_pair_of_openings Visible_as_one_opening,
  91  'F19-Pheromone glands' / Unbranched_tubes Branched_tubes Absent,
  92  'F20-Length of glands' / 'Short,_less_than_or_=_length_of_7th_sternite' 'Long,_greater_than_length_of_7th_sternite' Absent,
  93  'F21-Length of ductus bursa' / 'Short,_less_than_or_=_length_of_7th_sternite' 'Long,_greater_than_length_of_7th_sternite',
  94  'F22-Condition of ductus bursa' / Sclerotized_proximal_to_OB_and_membranous_closer_to_CB Completely_sclerotized,
  95  'F23-Ductus bursa with ornamentation' / Absent Present,
  96  'F24-Condition of corpus bursa' / Membranous Sclerotized_where_ductus_bursa_enters,
  97  'F25-Signa' / Absent Present,
  98  'F26-Signum type' / Absent Elliptical_scobinate_patches_with_inward_projecting_teeth Disc_shaped,
  99  'F27-Signa number' / Single_signum Two_signa Four_signa,
  100  'F28-Corpus bursa with secondary ornamentation' / Unmodified Scobinate_patch_surrounding_signum,
  101  'F29-Appendix bursa' / Absent Present,
  102  'F30-Location of appendix bursa' / Absent Attached_to_corpus_bursa Attached_to_ductus_bursa,
  103  'F31-Condition of appendix bursa' / Membranous Sclerotized Absent,
  104  'F32-Origin of ductus seminalis' / Arising_from_ductus_bursa Arising_from_corpus_bursa Arising_from__appendix_bursa Arising_from_duct_connecting_corpus_bursa_and_appendix_bursa Arising_from_duct_connecting_ductus_bursa_and_appendix_bursa,
  105  'F33-Position of Appendix bursa' / Ventral Dorsal
		;

MATRIX
[                                                       10        20        30        40        50        60        70        80        90        100   ]
[                                                       .         .         .         .         .         .         .         .         .         .     ]

'Argina astrea MD65/MD151'                     110010101011100010101110000100101102101111012220000002222000314120251000110003101111100001000010121012021
'Argina leonina MD64/MD153'                    110010100011100010001110000101011002101112114220000002212000415000000000110003101011101001001000111012020
'Mangina argus MD66/MD152'                     110???1010111000101011100001010001011012001110000001210220000110000000001111031011111000010110001110120?0
'Pitasila abraxoides 4031/2097'                ????????????????????????????11000?22101111010000100121020000213000000000????????????1000?????1001111?????
'Pitasila leucospilota 4066/4067'              ????????????????????????????01000?2210111101000010012112100001100000000011111?101?1?10000101?1001110?????
'Pitasila selecta 4052/4068'                   ????????????????????????????0?010?2210111101000010012112100021300000000011111?10101010000???10001111?????
Utetheisa_amhara_4275                         ???????0101110000???????????0?01011210111101100010112102100021112023100??????????????????????????????????
'Utetheisa bella 213/295'                    110010101011200000101110000101110101101112010000101120021100112000000121001113101011000002011000111012020
'Utetheisa clareae 2430/1597'                 ???????0101110001???????????????011210111101100010112102000021312013100100??1?1011101000010?10001110?????
Utetheisa_devriesi_1965                       0200001010111000001111100000000?010210111101211010112102101021312022300100111310101110000?111010111012020
'Utetheisa dorsifumata MD317/MD318'            1?00?01010111000101011100001010100221011110110101011211210002131203310010011131011111000????10101110?????
'Utetheisa elata MD219/MD272'                  111010101011100010101110000101110101101111012110101111020000213120131111001113101111100002111010111012021
'Utetheisa galapagensis MD259/MD296'         020000101011100000001110000000000002101111012110101121021010213120223001001113101111100002011000121012020
'Utetheisa lotrix MD214/MD267'                110000101011100000101110000101011012101111013000101120120110011113022001001111101111100002011110111012020
Utetheisa_lotrix_stigmata_1534                ???????0001110000???????????0?1?1112101111013000101121120110011112022001?????????????????????????????????
'Utetheisa lactea MD315/MD316'               1100?0101111100010101110000201110022101111012110101111021000113120111121001113101011100002011000111012021
'Utetheisa lactea MD368/MD400'                 110000101111100010101110000101110022101111012110101111021000113120111121001113101011100002011000111012021
'Utetheisa ornatrix MD215/MD339'             11?01010101110000010111000010?110101101112010000101120021100112000000121001113101011100002011000111012020
'Utetheisa perryi MD413/MD415'                 0200001000112000000111100000000100??101111012010101111021000213120131001001113101111100002011000121012020
'Utetheisa pulchella MD246/MD355'            110000100011100100101110000001110022101111012110101111021010213120121001001113101111100002011010111012021
'Utetheisa pulchella bicolor MD315/MD314'    1100?0100011100000001110000101011022111111012110101111021010213120121001001113101111100002011010121012021
'Utetheisa pulchelloides MD212/MD270'        11?000101011100010101110000001010022101111011010101121021000111120211001001113101011000002011010111012021
Utetheisa_pulchelloides_aphanis_1530          ???????0101110001???????????0?0?012?101111011010101121121000112120211001?????????????????????????????????
Utetheisa_pulchelloides_marshallorum_1524     ???????0101110001???????????0??00122101111011010101120021000112120211001?????????????????????????????????
Utetheisa_pulchelloides_vaga_1529             ???????0101110001???????????0????1??101111011010101121121000112120211001?????????????????????????????????
'Utetheisa salomonis MD307/MD309'            110000101001100000101110000000010022101111011010101121021000211120241001001113101111100001001010121012021
'Utetheisa semara MD311/MD312'                1100?0100011100000001110001200010032101111015000101120121110011120223001001111101111100002011100121012020
Atasca_pellex_MD260                          0?0000101?1110001???????????01101102111112111000100002220000111000000000?????????????????????????????????
'Haploa clymene MD19/MD20'                    1100101010112000001021200000010100021112000110001000022200003140000000001100131011111100010010001220110?2
'Euplagia quadripunctaria MD42/MD43'          1100001110111000001021100000010101021111120100011000022220004131201510000000131011111100010011001221110?2
'Tyria jacobaeae MD178/MD179'                1100101010112000001021200000000101421111120100001000022122003151201610000000031011111100010111001210110?2
;
END;


BEGIN ASSUMPTIONS;
	OPTIONS  DEFTYPE=unord PolyTcount=MINSTEPS ;
	WTSET * UNTITLED  = 1.00: 1-38 43-51 56-61 72 77-96 100 104, 0.50: 39-42 62-63 70-71 73-76, 0.25: 52-55 101-103 105, 0.16: 64-69, 0.33: 97-99;
END;


BEGIN TREES; 


	TREE  * UNTITLED =  [&R] ('Argina astrea MD65/MD151','Argina leonina MD64/MD153',(('Haploa clymene MD19/MD20',('Euplagia quadripunctaria MD42/MD43','Tyria jacobaeae MD178/MD179')),(('Mangina argus MD66/MD152',Atasca_pellex_MD260),(('Pitasila selecta 4052/4068',('Pitasila abraxoides 4031/2097','Pitasila leucospilota 4066/4067')),((Utetheisa_amhara_4275,'Utetheisa clareae 2430/1597'),((('Utetheisa lactea MD368/MD400',('Utetheisa elata MD219/MD272','Utetheisa ornatrix MD215/MD339')),(Utetheisa_pulchelloides_marshallorum_1524,(Utetheisa_pulchelloides_aphanis_1530,Utetheisa_pulchelloides_vaga_1529))),('Utetheisa dorsifumata MD317/MD318',('Utetheisa pulchelloides MD212/MD270',('Utetheisa salomonis MD307/MD309',(('Utetheisa semara MD311/MD312',('Utetheisa lotrix MD214/MD267',Utetheisa_lotrix_stigmata_1534)),(('Utetheisa pulchella MD246/MD355','Utetheisa pulchella bicolor MD315/MD314'),('Utetheisa perryi MD413/MD415',(Utetheisa_devriesi_1965,'Utetheisa galapagensis MD259/MD296')))))))))))));

END;


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