#NEXUS [ File saved by NDE version 0.5.0, Thu Apr 02 10:32:30 2009 ] BEGIN TAXA; DIMENSIONS NTAX=29; TAXLABELS 'Ero tuberculata ' 'Mimetus fernandi ' 'Mimetus syllepsicus ' 'Phobetinus sagittifer ' 'Australomimetus annulipes ' 'Australomimetus aurioculatus' 'Australomimetus burnetti ' 'Australomimetus catulli' 'Australomimetus daviesianus' 'Australomimetus diabolicus' 'Australomimetus djuka' 'Australomimetus hannemanni' 'Australomimetus hartleyensis ' 'Australomimetus hirsutus ' 'Australomimetus japonicus' 'Australomimetus maculosus ' 'Australomimetus mendax' 'Australomimetus miniatus ' 'Australomimetus sp. nov. "New Caledonia"' 'Australomimetus nasoi' 'Australomimetus pseudomaculosus ' 'Australomimetus robustus' 'Australomimetus sennio' 'Australomimetus sp. nov. "New Zealand"' 'Australomimetus spinosus ' 'Australomimetus stephanieae' 'Australomimetus sydneyensis ' 'Australomimetus tasmaniensis ' 'Australomimetus triangulosus' ; ENDBLOCK; BEGIN CHARACTERS; DIMENSIONS NCHAR=87; FORMAT DATATYPE=STANDARD MISSING=? GAP=- SYMBOLS="0123"; CHARLABELS [1] 'Overall body size' [2] 'Overall body shape' [3] 'Colouration of body integument (in alcohol)' [4] 'Maculation of the leg integument (in alcohol)' [5] 'Opisthosoma: shape' [6] 'Opisthosoma: dorsal setation' [7] 'Opishtosoma: with longitudinal whitish line on dark ground (ventral view)' [8] 'Opishtosoma: with distomedial subtriangular patch of creamy white colour (dorsal view)' [9] 'Opisthosoma: development of distomedial colour patch (dorssal view)' [10] 'Opishtosomal (dorsal view): distomedial colour patch present in juveniles' [11] 'Opishtosoma: distomedial colour patch present in juveniles (in alcohol)' [12] 'Spinnerets: shape of the cylindriform spigots' [13] 'Paturon: Numer of true teeth at distal interior margin' [14] 'Paturon: colouration' [15] 'Clypeus: setation' [16] 'Legs: formula' [17] 'Legs: femora I & II with a longitudinal row of short conical spines ' [18] 'Legs: extension of first row of spinules' [19] 'Carapace: extension of clypeus' [20] 'Carapace: colouration of cephalic region' [21] 'Carapace: with mediolateral brown colour spots ' [22] 'Carapace: development of mediolateral spots' [23] 'Carapace: shape of mediolateral colour spots' [24] 'Carapace: margin framed by broad marginal lines' [25] 'Carapace: anterior prolongation of pars chephalica' [26] 'Male pedipalp: functional position in which the palp is carried' [27] 'Ratio of the male pedipalpal patella/tibia' [28] 'Total numer of tibial trichobothria on the male pedipalpal tibia' [29] 'Total number of spines on the male pedipalpal patella' [30] 'Sclerites of the embolic division ' [31] 'Male pedipalp: shape of sclerites of the embolic division ' [32] 'Presence of a huge, simple sclerite in distomedial position (DMS) on the male pedipalp' [33] 'DMS of male pedipalp conical in lateral view' [34] 'DMS of male pedipalp diagonal in lateral view' [35] 'DMS of the male pedipalp hooked in lateral view' [36] 'Direction of tip of DMS in lateral view' [37] 'Concavity of tip of DMS in lateral view' [38] 'DMS with additional sclerites' [39] 'DMS with additional sclerotized plate ("SP") in subapical position' [40] 'Hooked DMS with broad apex' [41] 'Hooked DMS with pointed apex' [42] 'Hooked DMS sickle-shaped in frontolateral view' [43] 'Presence of S1 in the male pedipalp' [44] 'Presence of S2 in the male pedipalp' [45] 'Presence of S3 in the male pedipalp' [46] 'Shape of S3 in the male pedipalp' [47] 'Presence of S4 in the male pedipalp' [48] 'Presence of S5 in the male pedipalp' [49] 'Presence of a pedipalpal sclerite in medioectal position (MES)' [50] 'MES with additional cusps or cones' [51] 'Tegulum with distobasal sclerite or cone' [52] 'Basodistal sclerite plate-shaped and slightly elevated' [53] 'Basodistal sclerite as an elevated cone' [54] 'Tegulum in lateral view' [55] 'Size of subtegulum in relation to tegulum' [56] 'Medial curvature of tegular sperm-duct' [57] 'Male pedipalp: direction of embolus' [58] 'Presence of an EOS' [59] 'EOS: present and of very conspicuous shape' [60] 'EOS: present but inconspicuous' [61] 'Shape of cymbium' [62] 'Cymbium with a terminal sclerotization ("vexillum")' [63] 'Cymbium with a sclerotized cone in distoectal position' [64] 'Cymbium with conspicuous retrolateral sclerotized projection ("shovel")' [65] 'Cymbium with a short retrolateral sclerotized velum' [66] 'Cymbium with row of four slender and solitary spines ' [67] 'Cymbium retrolateral with three or four very thick setae' [68] 'Cymbium with megaspine(s) in subbasal position' [69] 'Size of paracymbium ' [70] 'Shape of paracymbium' [71] 'Orientation of elongate paracymbium ' [72] 'Basal width of elongate paracymbium' [73] 'Paracymbium with basal sclerotization or lobe' [74] 'Paracymbium with medial sclerotization or lobe' [75] 'Paracymbium with distal notch or broadened distally' [76] 'Paracymbium with velar medial lobe, projecting downwards.' [77] 'Shape of epigyne' [78] 'Epigyne with a distomedial "scape"' [79] 'Epigynal "scape" greatly exceeding distal wall of epigyne' [80] 'Epigynal "scape" with distal notch' [81] 'Copulatory openings conspicuously sclerotized' [82] 'Overall scerotization of epigyne' [83] 'Epigyne: with "pit" in posterior¨position' [84] 'Shape of receptacula' [85] 'Number of lobes of the receptacula' [86] 'Recepatacula: basally broadened or extended' [87] 'Orientation of receptacula' ; STATELABELS 1 'small (< 7 mm) ' 'medium (7-15mm) ' 'large (> 15 mm)', 2 'slender spiders with long legs in relation to carapace' 'robust spiders with short legs in relation to carapace', 3 'entirely brown or speckled' 'entirely pale', 4 'absent or inconspicuous' 'prominent and very strong', 5 'globular or oval' triangular, 6 'not dense, setae fine' 'dense, setae robust', 7 absent present, 8 absent present, 9 'not conspicuous or interspersed' 'conspicuous, not interspersed', 10 'absent or not visible' present, 11 'absent or not visible' 'present ', 12 'rotund, enlarged and incised' 'rotund, enlarged and smooth ', 13 two one absent, 14 'uniformily chestnut brown' 'uniformily pale' 'proximally pale with sharply demarcated darkened distal margin' 'uniformily pale but with proximal brown patch', 15 '5 setae, inner setae conspicuously smaller than outer ones' '5 setae, inner ones +/- as large as outer ones', 16 'I II IV III ' 'I IV II III', 17 absent 'present ', 18 'extending < half of coxa I' 'extending > half of coxae I', 19 'less than +/- AME tubercle diameter' 'about or more than +/- AME tubercle diameter', 20 'consisting of three or five median bands forming united triangular figure reaching fovea' 'consisting of three longitudinal bands with only median band reaching fovea' 'colouration absent or reduced', 21 absent present, 22 'prominent and conspicuous' 'not promiment and inconspicuous', 23 patches stripes 'fused, not individually discernable', 24 absent present, 25 'not or slightly prolonged' prolonged 'strongly prolonged', 26 ectal lateral, 27 'tibia < two times length of patella' 'tibia < three times length of patella' 'tibia > three times length of patella', 28 four six five three, 29 'one spine' 'two spines' 'three spines', 30 'complex and at least partly velar' 'relatively simple and heavily sclerotized', 31 'folded and divided in unexpanded pedipalp ' 'not folded and simple in unexpanded palp', 32 absent present, 33 'not conical' 'conical ', 34 'not diagonal' '(almost) diagonal', 35 'not hooked, not raised apically' 'hooked and raised apically', 36 'not diagonal ' diagonal, 37 'not concave' concave, 38 'without additional sclerites' 'with additional sclerites', 39 absent present, 40 'not broad' 'broad and massive', 41 'not pointed' pointed, 42 'not sickle-shaped' 'sickle-shaped', 43 absent present, 44 absent present, 45 absent present, 46 inconspicuous conspicuous, 47 absent present, 48 absent present, 49 absent present, 50 absent present, 51 absent present, 52 'inconspicuous or almost invisible' 'plate-shaped and slightly elevated', 53 'inconspicuous or almost invisible' 'developed as an elevated cone', 54 'not flattened or enlarged / not exceeding cymbium' 'flattened or enlarged /exceeding cymbium', 55 '< 2/3 of tegulum' '> 2/3 of tegulum', 56 'almost straight' 'slightly curved' 'strongly curved or looped', 57 distoectal medioectal, 58 'absent or invisible' present, 59 'not exceeding embolic origin' 'very conspicuous and exceeding embolic origin', 60 'present but not exceeding embolic origin' 'very small, hardly visible', 61 'cup-shaped' slender, 62 absent present, 63 absent present, 64 absent present, 65 absent present, 66 absent present, 67 absent present, 68 absent present, 69 'pronounced and well defined' 'small, may be incorporated into cymbium', 70 'shoveled or semi-square' 'elongate, distinctively longer than broad', 71 'paracymbium not bend downwards, rather straight without medial curvature' 'paracymbium proximally projecting downwards, medially curved and distally bent towards the interior (structure strongly concave from above)' 'not as above', 72 'rather slender, < double width of median part' 'broad, > double width of median part', 73 absent present, 74 absent present, 75 'notch absent' 'notch present', 76 absent present, 77 'triangular plate with transverse ridges, genital opening(s) inconspicuous' 'consisting of sclerotized medial plate and a distal plate' 'consisting of a single, huge sclerotized opening', 78 absent present, 79 'not exceeding distal wall of epigyne' 'greatly exceeding distal wall of epigyne', 80 absent present, 81 'not cospicuously scleroztized' 'conspicuously sclerotized', 82 'weakly sclerotized' 'strong sclerotization', 83 absent present, 84 globular 'oval or reniform' multilobed, 85 one more, 86 'not broadened' broadened, 87 'situated directly on basal plate with short genital ducts' 'space between receptacula and basal plate +/- diameter of receptacula', ; MATRIX 'Ero tuberculata ' 0100010000 0010110-11 0--1001301 1000000000 0000000000 0000100000 1000000001 0110101000 1100001 'Mimetus fernandi ' 0000010000 0013101000 1100111010 0000000000 0001111100 0000000000 0101000010 0-00000000 0012100 'Mimetus syllepsicus ' 0000010000 0013101000 1100110000 0000000000 0011101100 0000000000 0111000000 0-00000000 0012100 'Phobetinus sagittifer ' 0000110000 0?11?01000 1020110001 -000000000 0000000000 00001-1000 000001001- --00002000 0112100 'Australomimetus annulipes ' 2101010111 0110001000 10011????? ?????????? ?????????? ?????????? ?????????? ??????1000 110?001 'Australomimetus aurioculatus' 0010000000 1111001000 1000102011 1100101000 0000000000 0000020000 1000010001 0011001000 0100001 'Australomimetus burnetti ' 1101010111 0110001000 1000100221 1100101001 0000000000 0000010101 1000000001 1101111001 1101001 'Australomimetus catulli' 1101010101 0110001000 1000101121 1100100101 0000000011 0000011110 1000110001 111100???? ??????? 'Australomimetus daviesianus' 1101010101 0100001000 1010101121 1100100010 1100000000 0000021000 1000001001 0001101110 0101000 'Australomimetus diabolicus' 1010000000 0112001000 1000101021 1100100100 1000000010 0000121000 100000101- 0010101000 0101010 'Australomimetus djuka' 1101011000 0100001010 1001101021 1100110000 0000000000 0000110000 1000000001 0000001100 0101000 'Australomimetus hannemanni' 1101010101 0110001000 1101101221 1100101001 0000000000 0000010101 1000100001 110110???? ??????? 'Australomimetus hartleyensis ' 1101010101 0110001000 1001101221 1100101001 0000000000 1100010101 1000000001 1110111111 1101010 'Australomimetus hirsutus ' 0010100000 1111001000 1000101021 1100100000 1000000000 0000010000 1000000001 1010101000 0100011 'Australomimetus japonicus' 1010010000 0?11001000 1000101?21 1100100000 0000000000 0000120101 1000000001 0000101000 0101001 'Australomimetus maculosus ' 2101010001 0110001100 1000201121 1100100110 1100000010 0000111000 1000001001 0000001100 0101000 'Australomimetus mendax' 1101010101 0110001000 1100100211 1100100100 1000000011 1010011101 1000000001 0010101100 1101000 'Australomimetus miniatus ' 0010100000 1111001000 1000101021 1100100100 0000000011 0000110000 1000000001 2100001000 0101011 'Australomimetus sp. nov. "New Caledonia"' 0010000001 0110001000 1000102011 1100101000 0000000000 0000111000 1000000001 0011101000 1101000 'Australomimetus nasoi' 2101010101 0110001002 1000110221 1100100000 1000000000 1100110101 1000000001 101000???? ??????? 'Australomimetus pseudomaculosus ' 2101010101 0110001000 1000101121 1100100110 1100000010 0000111000 1000001001 0000001000 0101000 'Australomimetus robustus' 2101010111 0110001000 10011????? ?????????? ?????????? ?????????? ?????????? ??????1111 1101010 'Australomimetus sennio' 110101000? 0?10001010 0111001221 1111000100 0000000011 0001021110 100000011- --1010???? ???0000 'Australomimetus sp. nov. "New Zealand"' 110101000? 0?10001010 1110101021 1100101001 0000000000 0000021101 1000000001 0010101000 0100010 'Australomimetus spinosus ' 1101010000 0112001000 1110101221 1111000110 0000000011 0001021000 1000000011 000110100- 0100000 'Australomimetus stephanieae' 1101010000 0112001000 1111101021 1111000110 0000000011 0001021000 100001011- --01001000 0100000 'Australomimetus sydneyensis ' 1101010101 0100001000 1110100211 1100100100 1000000011 1010021101 1000000001 0010101100 1101000 'Australomimetus tasmaniensis ' 0100011000 0100011010 0--1000301 1100110000 0000000000 0000110000 1000000001 0001001100 0101000 'Australomimetus triangulosus' 0010100000 1111001000 11001????? ?????????? ?????????? ?????????? ?????????? ??????1000 0101000 ; ENDBLOCK; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE=UNORD POLYTCOUNT=MINSTEPS; ENDBLOCK; BEGIN NOTES; [Taxon comments] [Character comments] TEXT CHARACTER=15 TEXT='The Australian type consists of two pairs of sutural setae, with the inner ones smaller than the outer ones. More proximal, in between the AME, another setae is found, that is apparently absent in American and African Mimetidae.'; TEXT CHARACTER=20 TEXT='bleeched in tpye specimens of A. miniatus but to be seen as slight darkenings in full light'; TEXT CHARACTER=21 TEXT='fused in Phobetinus and A. hirsutus'; TEXT CHARACTER=24 TEXT='A carapace framed by a dark lateral line is typical for most Ero species. Homoplasic features are found in Ero tasmaniensis, Australomimetus 1 and the New Zealand species.'; TEXT CHARACTER=25 TEXT='A strongly elevated pars cephalica is typical for Australomimetus maculosus and Australomimetus pseudomaculosus. The pars cephalica is relatively inconspicuous in Ero, Ero tasamaniensis and Mimetus sennio.'; TEXT CHARACTER=26 TEXT='Mimetidae with a "shovel" rotate the palps to a functionally lateral position. Australomimetus and Ero carry the palps functionally ectal. Some dead Australomimetus show the lateral condition, but this might be artificial because investigations from the living specimens prove the opposite.'; TEXT CHARACTER=27 TEXT='Although the character is somewhat subjective, it seems to be useful since most American Mimetus and Phobetinus share very short tibiae whereas Ero and Australomimetus have medium sized tibiae. The New Caledonian species and Mimetus aurioculatus share the synapomorphic character of very long tibiae.'; TEXT CHARACTER=28 TEXT='Subject to homoplasies, this feature is consistent within one species but of limited phylogenetic value.'; TEXT CHARACTER=30 TEXT='The distal sclerites referred to as terminal apophysis (Shear, 1981) or median apophysis (Heimer, 1986) are undevided in Mimetus syllepsicus and Mimetus fernandi. These species share the synapomorphic condition of these sclerites in equal position and equal shape (P1-P5), which are sometimes velar (P2) or fragile (P3, P5). None of these features is present in Australomimetus, where the distal sclerites of the male palp are always robust, heavily sclerotized and close to each other. Similar, but surely homoplasic conditionsd are present in Ero and Phobetinus. In Phobetinus a secondary simplification is reasonary since other features of the male palp (f. ex. the shovel or the paracymbium) are also subject to reduction and simpification.'; TEXT CHARACTER=39 TEXT='greatly exceeding DMS in A. daviesanus'; TEXT CHARACTER=59 TEXT='not comspicuous'; TEXT CHARACTER=62 TEXT='Synapomorph for North-American Mimetus, some African and Asian Mimetus. Neither present in Australomimetus nor Ero or Phobetinus.'; [Character state comments] [Attribute comments] [Taxon pictures] [Character pictures] [Character state pictures] [Attribute pictures] ENDBLOCK;