#NEXUS [NEXUS FILE GENERATED BY WINCLADA (Copyright 2002 K.C. Nixon); THIS DOES NOT CONSTITUTE ENDORSEMENT OF THIS FORMAT, NOR GUARANTEE ACCURACY] BEGIN DATA; DIMENSIONS NTAX=50 NCHAR=200; FORMAT MISSING=? SYMBOLS= " 0 1 2 3 4 5 6 7 8 9"; OPTIONS MSTAXA=POLYMORPH ; CHARLABELS [1] 'wing_margin_proximal_to_MP-c' [2] length_of_MP [3] Pigmentation_wing_apex [4] shape_of_wing_RA [5] posterior_margin_of_wing_at_MPc [6] wing_long_setae_at_margin [7] wing_jugal_vein [8] posterior_margin_of_wing_in_anal_region [9] wing_AA_vein [10] wing_MP_anteriorly [11] 'RP-RA_veins' [12] wing_AA_vein_shape [13] wing_jugal_vein_and_AP [14] wing_Z_vein [15] wing_Z_vein_at_posterior_margin [16] wing_setae_costal_margin [17] wing_anal_region_reduced [18] wing_MP_+_CuA_separation [19] wing_costal_margin_at_base [20] anterior_median_process_apically_dorsal_view [21] lateral_combs [22] anterior_median_ventral_process [23] anterior_median_ventral_process [24] posterior_median_tormal_process [25] lateral_tormal_process_proximally [26] lateral_tormal_process_length [27] apical_margin_of_distal_epipharynx [28] apical_margin_of_distal_epipharynx_excluding_projection [29] apex_of_anterior_median_dorsal_process [30] proximal_epipharynx_near_trans._median_suture [31] anterior_median_process_at_base [32] opposite_to_anterior_median_process_at_base_on_ventral_surface [33] apex_of_anterior_median_dorsal_process_assoc._setae [34] apical_margin_of_distal_epipharynx [35] shape_of_cavity_on_dorsal_side_of_proximal_portion_of_epiphar. [36] cavity_on_dorsal_side_of_proximal_portion_of_epi. [37] anterior_median_process_apex [38] ventral_surface_setae [39] lateral_tormal_arms [40] apex_of_ventral_articulatory_sclerite_of_galea [41] galea_shape [42] proximal_inner_strut_of_lacinia [43] apical_palpomere_color [44] distal_half_of_ventral_articulatory_scle._of_galea [45] distal_half_of_vent._artic._sclerite_at_apex [46] lateral_sclerite_notch_where_palp_inserts [47] accessory_internal_sclerite [48] accessory_internal_sclerite_proximal_tip_anteriorly [49] outer_tooth_of_lacinial_articulation_sclerite [50] ventral_artic._sclerite_dorsal_view [51] number_of_channels_associated_with_galea [52] 'ventral_artic._sclerite_anterio-laterally' [53] number_of_palpomeres [54] second_palpomere_size_compared_to_first [55] third_palpomere_compared_to_second [56] attachment_point_of_third_palpomere [57] shpae_of_first_palpomere [58] first_palpomere_laterally_expansion [59] mentum_apical_edge_medially [60] mentum_apical_margin_near_lateral_edge [61] glossa_internal_strut [62] mentum_shape [63] mentum_posterior_lateral_edge [64] paraglossae [65] paraglossal_apex [66] paraglossal_strut_in_lateral_view [67] paraglossal_strut_at_proximal_apex [68] palpomere_strut_on_recurved_portion_in_lateral_view [69] paraglossal_lobes [70] shape_of_anterior_declivous_region_of_mentum [71] between_paraglossae_at_their_base [72] most_proximal_apodemes [73] apodeme_adjacent_to_the_paraglossal_apodeme_in_dorsal_view [74] posterior_arm_of_apodeme_adjacent_to_the_paraglossal_apodeme [75] proximal_transverse_bridge_of_posterior_labial_struts [76] proximal_transverse_bridge_of_posterior_labial_struts [77] longitudinal_arms_adjacent_to_distal_trans._bridge [78] distal_transverse_bridge_in_ventral_view [79] 'dorso-ventral_arm_from_distal_bridge' [80] most_proximal_apodeme [81] cervical_scle._apex [82] cervical_scle._lateral_margin [83] cervical_setal_brush_position [84] length_of_setal_brush_located_mesally_near_apex [85] cervical_lobe_at_or_near_apex [86] cervical_orientation_of_apodeme_near_base [87] cervical_apex_of_muscle_attachment_point_shape [88] sclerotized_area_extending_towards_mandible_apex_medially [89] apex_of_sclerotized_area_near_inner_lateral_margin [90] 'sclerotized_area_between_long._carina_and_in._lat._edge,_sensilla' [91] sclerotized_area_between_long_car._and_in._lat._edge_setae [92] distinct_longitudinal_carina_near_lateral_edge_at_middle [93] apical_reach_of_mesal_comb [94] receptacle_shape_at_proximal_margin [95] 'receptacle_towards_brush/comb_side' [96] antennal_scape_setal_distribution_dorsally [97] antennal_scape_setal_row_nearest [98] antennal_scape_setal_row [99] antennal_scape_setal_row_extending [100] scape_elbow [101] seventh_antennomere [102] apical_antennomere [103] seventh_and_eigth_ant. [104] abdom._spiracle_position [105] abdom._spiracle_width [106] prescutum_prephragma [107] prescutum_anteriorly_trans._plate [108] prescutum_anteriorly_trans._plate_projection_shape [109] prescutum_anteriorly_tran._plate_anterior_edge [110] prescutum_anteriorly_trans._plate_apodeme [111] 'scutum-scutellum_lateral_edge_' [112] scutellum_apex [113] 'scutum-scutellum_levels' [114] 'prescutum_dorso-ventral_trans._plate' [115] axillary_cord [116] prephragmal_arm [117] scutellum_posterior_margin [118] prescutum_posterior_margin [119] prescutal_tongue [120] scutellum_anterior_margin [121] mediophragma_middle [122] mediophragma_cleft_shape [123] scutellum_posterior_margin_setation [124] scutellum_width_apically_compared_to_length [125] alar_ridge_of_scutellum [126] 'pyg._anterio-medial_groove_edges' [127] 'pyg._anterio-medial_groove_shape' [128] 'pyg._anterio-medial_groove_length' [129] pyg._transverse_ridge [130] 'pyg._shape_of_dorso-lateral_edge' [131] propygidium_at_middle [132] pygidium_pigmentation [133] propygidium_length [134] paramere_apex [135] paramere_position [136] dorsal_plate [137] ventral_plate [138] spiculum_gastrale [139] clypeal_teeth_number [140] genal_development_laterally_ [141] genal_lateral_edge [142] 'gula_apical_1/2' [143] gula_lateral_edge [144] gula_proximal_margin [145] clypeus_ventrally [146] frons_projection [147] eye_canthus [148] epipleura_notch [149] elytral_striae [150] elytral_surface [151] first_ventrite_visibility [152] sixth_ventrite_visibility [153] sixth_ventrite_length_at_middle [154] first_ventrite_visible_projection [155] first_ventrite_hidden_projection [156] third_ventrite_at_middle [157] elytra_pseudoepiplura [158] elytral_intervals_dorsally [159] tibial_teeth [160] tarsi_present [161] protibial_apex [162] tibial_carina [163] pronotal_dimple [164] profemoral_brush_setae [165] pronotum_lateral_edge [166] pronotal_surface [167] prothoracic_apodemes_large_plates [168] prothoracic_apodemes_horizontal_surface [169] proth._apo._simple [170] protibia_microteeth [171] proth._apodeme_with_large_anterior_amorphous_lobe [172] metasternum_posteriorly_with_depression [173] metasternum_with_anter._spine [174] mesosternum_with_post._edge [175] metatarsal_claws [176] mesotibial_apex_spines [177] 'first_-_fifth_metatarsomere_length' [178] 'first_-_second_metatarsal_length' [179] midcoxae_orientation [180] metatibial_expansion_towards_apex [181] distal_position_of_lateral_chitinous_line [182] shape_of_lateral_chitinous_line [183] form_of_posterior_area_of_furcal_arms [184] direction_of_furcal_arm_tip [185] furcal_tip_shape_apically [186] width_of_furcal_arm [187] chitinous_line_post._furcal_arms [188] form_of_midline [189] 'form_of_post-_lat._projection' [190] form_of_posterior_margin [191] shape_of_main_body_towards_post. [192] frontal_triangle_projection_directed [193] frontal_triangle_position [194] frontal_tip_shape [195] orientation_of_frontal_midline [196] frontal_midline_position_at_apex [197] shape_of_lateral_chitinous_projection [198] orientation_of_lateral_chitinous_projection [199] form_of_ventral_edge_post._of_lat._chit._projection [200] width_near_posterior_attachment ; STATELABELS 1 , 2 , 3 , 4 , 5 , 6 , 7 , 8 , 9 , 10 , 11 , 12 , 13 , 14 , 15 , 16 , 17 , 18 , 19 , 20 , 21 , 22 broad_entire_length not, 23 broad_base_relatively_long short, 24 , 25 , 26 , 27 , 28 , 29 , 30 , 31 , 32 , 33 , 34 , 35 , 36 , 37 , 38 , 39 , 40 , 41 , 42 , 43 , 44 , 45 , 46 , 47 , 48 , 49 , 50 , 51 , 52 , 53 , 54 , 55 , 56 , 57 , 58 , 59 , 60 , 61 , 62 , 63 , 64 , 65 , 66 , 67 , 68 , 69 , 70 , 71 , 72 , 73 , 74 , 75 , 76 , 77 , 78 , 79 , 80 , 81 , 82 , 83 , 84 , 85 , 86 , 87 , 88 , 89 , 90 , 91 , 92 , 93 , 94 , 95 , 96 , 97 , 98 , 99 , 100 , 101 , 102 , 103 , 104 , 105 , 106 , 107 present absent, 108 , 109 , 110 , 111 , 112 , 113 , 114 , 115 , 116 , 117 , 118 , 119 , 120 , 121 , 122 , 123 , 124 , 125 , 126 , 127 , 128 , 129 , 130 , 131 , 132 , 133 , 134 , 135 , 136 , 137 , 138 , 139 , 140 , 141 , 142 , 143 , 144 , 145 , 146 , 147 , 148 , 149 , 150 , 151 , 152 , 153 , 154 , 155 , 156 , 157 , 158 , 159 , 160 , 161 , 162 , 163 , 164 , 165 , 166 , 167 , 168 , 169 , 170 , 171 , 172 , 173 , 174 , 175 , 176 , 177 , 178 , 179 , 180 , 181 , 182 , 183 , 184 , 185 , 186 , 187 , 188 , 189 , 190 , 191 , 192 , 193 , 194 , 195 , 196 , 197 , 198 , 199 , 200 , ; MATRIX Cyptochirus 001010001111-00100010111110010010001100001010101013200--1010211101110001010001101011111001100010110011003101211010112010101002--0121011011300010100101001100130001?0011010121101022101113131122002120120 Scaptocnemis 101010001111100100010111111011110001100001010101013201--2010211101110011110000100011111101100010110011003101211010112010101002--0101101??110001110011100110102000100001010121101022111123101122002120130 Euoniticellus 010110111111-00100010100101111111001100101110101013211002010201101110011020?011?1011111111100011110011003001511011112010101002--21-0-1101110001112010100110002000100001010121101022201113110??2012101130 Oniticellus 000110111111-0010000011?111110010101100101010101013200--2010211101110031010101101011111111100011110011003100521011112010101002--21-0-1101110001010010100121002000100001010121101022221403131111002221130 Drepanocerus 101010111111-00100010111110211010101100001010101013200--2011210101110011010101101011111011100011110011011101211110112010101000020101111011100011101101001100130001?0011110101101022001114101122002210130 Tiniocellus 001010011111-00100010111121411010101100001110101013200--1021211101110031110100111011111111100011101011023000511011112010101002--21-0-1101100001110010100121112000100001010121101022201213130122011120110 Liatongus 001010111111-00100010111121210110001100001010101013200--1010201101110001010101101011111111100010111011023101521011112010101002--21-0-1101100001010010100110002000100011110121101022201113100121002220010 Tragiscus 000110012111100100000111111010110101100101001001003100--2010201101110031010011101011111111100410111011003100521011112011101002--22-1-1101111001012010100110012000000011010121101022201223110011002221010 Neosisyphus 021110111101-00100110111321111110101100010010101002211101020010101110011000121111010110101100211101011112100501101103004001110220121001102100010101111001101011001000011011011011220011021110320022001?0 Anachalcos 0001110101010110001201000-130111110100011110010011311001323001000111001100111112001101111110131001001111110040010100101000111011020111100210010010110100121123110000001111121101111001323130101000100100 Copris 0001100101010110001200112-1811100100101111100011011110100-2111010111001101011001001111111110031021001111210021010011201400110001011110100211101012110110120111100100011110121101212211212111101000001100 Onitis 000201100121-10100121111101111110101101010111001000210002020200100110001140011010010111101010310110011103100311001112013110102--01011110070000101101010012102?011000001010111001012212223100112102120110 Garreta 00021000112011010212011110121110011000111000102-003110100-1121010111001101011111101011011110121021101111110131010111101000111101000110100340020210100000120101100000001111121100111001421000022001221220 Phanaeus 00010101011111010012011010190110001100101110001001301100212130010011130111011110001111111110021021101112110020110011221011010001022100100601001012010101110002100100011110120111011202323100112113120010 Canthon 000211111121-10101130101101411110111001011001001013111002031010000101311010120010010011111101211111011111100111101002010001212--020100110520021010110100120011100100001111121101111002223200101012210030 EurysternusV 100210112011---0101421100-151011210111101111002-013200--113001010111003122010210101011111110021011001110310011101111110310010002012101000201000010111100010001101100001101101101022002313131122101121121 Kheper 0001110001001100001211110-0411001101001011101001210210100-0101010001020103311111000-10111101120001001111010002110111101112110111013120100341100010010000120123010001101110021102110020404130111001120111 Dichotomius 000211000001111000120101101301000101001011100001011311102020210000111100000120100010111110100210201100121100200100012?1101011001010111100110001002110100121012100100011110101101211212000020101000200110 Onthophagus 0000100111011001010401110-101111010110111111010100321000102101010111001104002110011011101110001021001102210131210111301000121100121111100100001011010100121102000100011110121101021201123101111012021130 Metacatharsius 000111010101-110001200113102011001-2101011101011011010100-2101010011000101001000101111110101031121100011110030010111101200111001023110100211100012110100120112100100011110121101012201213130111010001100 Diastellopalpus 000000011111-0010102011112121111010110001110010101011101212010010110010114001110011011100110001021001102210131210111201002021100123111100100001012010100121002000100011110120101021212413100121002220110 Canthidium 000110111101-1100012000111181110010100001111012-210210201211110101110?0101??111?00101111101012110011101221004001011122120011100001111110-220001000110100120112100100001110121101012201121100101010200130 Glyphoderus -------------------2011011130111210100101100002-21?210123221010111110120112110001011111011100111201201120110101101?1---------1?1000110110231100202010100120013010100110110121101011000212100100021001210 Scarabaeus 0001110021001110001201110-111100000100101110102-210210100-0101010001020003011111000010111101120001101111010032110101?011?2110111010111100341120010011100120023010001101111121100110011300100111001100100 Oxysternon 0001010101111100001201112-1901000011001011100010013011102020300110111301110?????001011111110031101101111110020110011101001011001010110100601001002010101110002110000011110120111211201303100102000100110 EurysternusC 000210112011-001101401110-101011010110101111002-003200--101121110111001122010010101111111110001021001110310001101111210210110022011110000200000010111100010001101100001101121101022001323131132002000110 Epirinus 1001110101011100011201110-1612000102100011000001001110101011010100110011010111101011111001101?11111011111100121101101014001111200201101002111010101111002200121001?0001111121101111001323110110010000100 Pedaria 0002110100010110001001112-171110010110101111000100031110221101010111000101011110101011111110001110101111110012010111101400110002030111100210001110110110320110100110001110121101131201112111112000200100 Sulcophanaeus 0001110101111100001201100-1911000011001011100010012011102120310100111001110111100010111111100210011011111100201100112211010100010101011?0611001012010101120002100100011110120111111201023100101000100110 Xinidium 000101010001111000120111301611110101101110110101000110101221110101110011000011100010111111100311211000121100310101111014001110000201111002211???01110110220110000100001110121101212201310000101012011010 Bubas 001200100121-10100120111301611110101101110110101000210031220100101110001140010000010111011010210110011103100311101112013110102--010101100700001011010100121001011100011010111001022222021000032002000130 Anomiopus 000210110121-11110120111311911100001000011110?2-210201--2221110100110011010110000011111111110311211110111100300101112010001110000101112??230001010110100220011100110001111121101011301313100101002000120 Coptodactyla 0001010101010111101201110-170111010110001011000101021010122011010111000101011110001111111110031021000112110030010111201100111001010111100110000012110110120110110100011110121101211201210031101000200110 Scatimus 000111010101011000120111301311100101000011111011000210100-1101010011001101012110001111111111031???010012110031010111201100111010020110100220001101110100220112100100001110121101211201300101100012201210 Ateuchus 00011101010101000012001131170110010110001111111121121010101011010111000101011010001111111110031111011111110031010111001400111000010110100120001110110100120112102100001110121101011200323101101000200000 Heteronitis 001101100101-1010012111110111111010110111111000100121003203010010111010114000001001011101110021011001110210031110011101311010100000101100700001010010100121022011100011010111001022222323100032101100110 Coptorhina 000210112121-111001201110-140110010100111-11102-21?210100-0101010111000131?11112001111111010101001011012110020110111121100111011010111110420000212100110221110100100011110121101211201220011101002120220 Sisyphus 121110111111-00000110111311111110101100110010101011211101220010101110001010121111010110101100210111011112100501101103004001110220121001101100010101111001101021001?0001101101101222001422110132013101130 Anomiopsoides -------------------2011111130111210100101101002-213210123221010111110120012110011011111011100111201201120110101101?1---------????????01102411002010101001??013010100110110121101011000100030101021000010 Circellium -------------------201110-11010001010000111000012011110020110101001100010301???10010111011101?100100111111101?1101012011--11100101312010022110001001-100120113110110001110121101111010410011111220000211 Colobopterus 000010111101-2-1001501--0-12-2-1-1-2-110110100??010310101211010101110031320100030011111111110011010110121100221111112010001102--22-1-0110400001010110100000000100110001110121101013211220020101002000102 Amphistomus 100111000101011001120111??16100101011010111101010102100012200101011100310201111000111111011003102000111111004101011010100011100002111110?230011100111110111113100100001111131100112011221031032000200231 Lepanus 000111000001-1100012000132071111000110001111000?21121000101101010011001100012010001111101110031110101111110012010111001100111000021110100220001100110100220012101100001111121101101001414100101010200020 Ataenius 00001000110102-1011531-13106-0-1-101-0101111112-203310200-1101012111043130013000001111111110201001011012111061101111201000110011020110110401101110110110220010100110001110121101023101021011111112000002 Demarziella 00011000010101100011011132141010010010101111012-00021000201111010011000001011?11101111101110031120101011110061010111201100111000020110100120001000111110320110100110001110121101131202203101101000002020 Arachnodes 000111000001111000120011311811100100100011110101210210100-1121010011001102011010001111110110031120101111110051010110201100111000020110100420001100110110121112100100001111121101101011110000122023001230 Synapsis 00010001010101111112011111151100010100101110100020011110212121001111010013011111001011110101020001010111110030010001101211011001010110100221100012110100221113100000001111001101012122000010111001020100 Bdelyropsis 000110000101011000120111301711100101100111111001001110101011010100110011010111100010111110100012---11112110021010011001100111001030110110420001100110100221012102100001110121101211101213101110002120110 Cephalodesmius -------------------2010030181111110100101111112-003110100-1101010011000102??311?001011101100031021111112111---010-103?111-111110012100100640011010011100121113100100101111121101131010423100131021000110 Heliocopris 000100010101011000120111111511000101101111111111010111011220200001110?1110213110001111111101120001010111110000010001101011111011001110110200000011110100121113100000011110001101011221212100111000000110 ; ENDBLOCK; BEGIN ASSUMPTIONS; OPTIONS DEFTYPE=unord PolyTcount=MINSTEPS ; ENDBLOCK;