Plantations are the most common means of reforestation following stand-replacing wildfires. As wildfires continue to increase in size and severity as a result of fire suppression or climate change, establishment of plantations will likely also increase. Plantations’ structural characteristics, including dense, uniform spacing and abundant ladder fuels, present significant wildfire hazards. Large-scale fuels reduction techniques may be necessary to reduce potential fire behavior in plantations and to protect surrounding forests. In the present study, four different manipulations aimed at reducing potential fire behavior in a Sierra Nevada pine plantation are compared. The treatments include: mechanical shredding, or mastication, of understorey vegetation and small trees; mastication followed by prescribed fire; fire alone; and controls. Fire behavior modeling shows that mastication is detrimental whereas prescribed fire is effective in reducing potential fire behavior at moderate to extreme weather conditions. Predicted fire behavior was compared with actual values from the prescribed burns in an effort to explore the limitations of fire modeling. Fire behavior predictions were similar to field observations in the more structurally homogeneous stands, but differed greatly where mastication created forest openings and patchy fuels distributions. In contrast to natural stands, the homogeneity of pine plantations make the results of the present work applicable to other regions such as the south-eastern US, where similar fuels reduction techniques are used to increase fire-resistance and stand resilience.