References

Acha, E. M., Mianzan, H. W., Guerrero, R. A., Favero, M., and Bava, J. (2004). Marine fronts at the continental shelves of austral South America. Physical and ecological processes. Journal of Marine Systems 44, 83–105.
| CrossRef |

Burnham, K. P., and Anderson, D. R. (2002). ‘Model Selection and Multimodel Inference: A Practical Information–Theoretic Approach.’ (Springer–Verlag: New York.)

Coulson, J. C. (1968). Differences in the quality of birds nesting in the centre and on the edge of a colony. Nature 217, 478–479.
| CrossRef |

Coulson, J. C., and White, E. (1956). A study of colonies of the Kittiwake Rissa tridactyla (L.). Ibis 98, 63–79.
| CrossRef |

Coulson, J. C., and White, E. (1958). The effect of age on the breeding biology of the Kittiwake Rissa tridactyla. Ibis 100, 40–51.
| CrossRef |

Crawley, M. J. (2007). ‘The R Book.’ (Wiley: West Sussex, UK.)

Daunt, F., Wanless, S., Harris, M. P., and Monaghan, P. (1999). Experimental evidence that age-specific reproductive success is independent of environmental effects. Proceedings of the Royal Society of London. Series B. Biological Sciences 266, 1489–1493.
| CrossRef |

Daunt, F., Wanless, S., Harris, M. P., Money, L., and Monaghan, P. (2007). Older and wiser: improvements in breeding success are linked to better foraging performance in European Shags. Functional Ecology 21, 561–567.
| CrossRef |

Furness, R. W., and Monaghan, P. (1987). ‘Seabird Ecology.’ (Blackie: Glasgow.)

Lack, D. (1968). ‘Ecological Adaptations for Breeding in Birds.’ (Methuen: London.)

McCullagh, P., and Nelder, J. A. (1989). ‘Generalized Linear Models.’ (Chapman and Hall: New York.)

McNeil, R., and Léger, C. (1987). Nest-quality and reproductive success of early- and late-nesting Double-crested Cormorants. Wilson Bulletin 99, 262–267.

Moreno, J. (1998). The determination of seasonal declines in breeding success in seabirds. Etología 6, 17–31.

Nelson, J. B. (2005). ‘Pelicans, Cormorants and Their Relatives. The Pelecaniformes.’ (Oxford University Press: Oxford, UK.)

Perrins, C. M. (1970). The timing of birds’ breeding seasons. Ibis 112, 242–255.
| CrossRef |

Quintana, F., and Yorio, P. (1998). Kelp Gull predation on an Imperial Cormorant colony in Patagonia. Marine Ornithology 26, 84–85.

Quintana, F., Wilson, R., Dell’Arciprete, P., Shepard, E., and Gómez Laich, A. (2011). Women from Venus, men from Mars: inter-sex foraging differences in the Imperial Cormorant Phalacrocorax atriceps a colonial seabird. Oikos 120, 350–358.
| CrossRef |

R Development Core Team (2009). ‘R: A Language and Environment for Statistical Computing.’ (R Foundation for Statistical Computing: Vienna, Austria.) Available at http://www.R-project.org [Verified 14 May 2011].

Rivas, A. L., Dogliotti, A. I., and Gagliardini, D. A. (2006). Seasonal variability in satellite-measured surface chlorophyll in the Patagonian Shelf. Continental Shelf Research 26, 703–720.
| CrossRef |

Saether, B. E. (1990). Age–specific variation in reproductive performance of birds. In ‘Current Ornithology. Vol. 7’. (Ed. D. M. Power.) pp. 251–283. (Plenum Press: New York.)

Schreiber, E. A., and Burger, J. (2002). ‘Biology of Marine Birds.’ (CRC Press: Boca Ratón, FL.)

Shaw, P. (1986). Factors affecting the breeding performance of Antarctic Blue-eyed Shag Phalacrocorax atriceps. Ornis Scandinavica 17, 141–150.
| CrossRef |

Siegel-Causey, D., and Hunt, G. L. (1981). Colonial defense behavior in Double-crested and Pelagic Cormorants. Auk 98, 522–531.

Snow, B. K. (1960). The breeding biology of the Shag on the island of Lundy, Bristol Channel. Ibis 102, 554–575.
| CrossRef |

Staverees, L., Crawford, R. J. M., and Underhill, L. G. (2008). Factors influencing the breeding success of Cape Gannets Morus capensis at Malgas Island in 2002/2003. Ostrich 79, 67–72.
| CrossRef |

Svagelj, W. S. (2009). Breeding ecology of dimorphic seabirds in relation to parental investment and brood sex ratio theories. Ph.D. Thesis, University of Buenos Aires, Argentina. [In Spanish]

Sydeman, W. J., Penniman, J. F., Penniman, T. M., Pyle, P., and Ainley, D. G. (1991). Breeding performance in the Western Gull: effects of parental age, timing of breeding and year in relation to food availability. Journal of Animal Ecology 60, 135–149.
| CrossRef |

Velando, A., and Freire, J. (2001). How general is the central–periphery distribution among seabird colonies? Nest spatial pattern in the European Shag. Condor 103, 544–554.
| CrossRef |