
TY  - JOUR
A1  - Thayer, Margaret K. 
T1  - <emph type="2">Glypholoma</emph> larvae at last: phylogenetic implications for basal Staphylinidae? (Coleoptera : Staphylinidae : Glypholomatinae)
JF  - Invertebrate Systematics
JO  - Invert. Systematics
Y1  - 2000
VL  - 14
IS  - 6
SP  - 741
EP  - 754
KW  - 
UR  - https://doi.org/10.1071/IT00019 
N2  - Species of Glypholoma Jeannel, a genus previously moved to Staphylinidae from Silphidae, have adult synapomorphies of the Omaliine Group, a probably basal lineage of about 10 000 species. Adult characters have not definitively resolved the position of Glypholomatinae within the Omaliine Group, but suggest that it may be relatively basal. Larvae and pupae recently collected with adults of Glypholoma pustuliferum Jeannel are attributable to that species on the basis of that association, a good match of size and form between pupae and adults, known larval synapomorphies in Staphylinidae and background knowledge of the Chilean fauna; they are described and illustrated. Structural and chaetotaxic characters of these and larvae of 25 other Omaliine Group genera were subjected to phylogenetic analysis. The position of Glypholomatinae within the Omaliine Group remains uncertain, although it is weakly supported as forming a clade with Omaliinae and Microsilphinae. Monophyly of the subfamily Proteininae and its components and of the tribe Coryphiini of Omaliinae are more strongly supported. Resumen. Las especies de Glypholoma Jeannel, un g&#233;nero de Staphylinidae transferido previamente de Silphidae, presentan en estado adulto sinapomorfias con el Grupo Omaliine, un linaje probablamente basal constituido por alrededor de 10 000 especies. Los car&aacute;cteres de los adultos no han resuelto definitivamente la posici&oacute;n de Glypholomatinae dentro el Grupo Omaliine, pero sugieren que es relativamente basal. Se describe e ilustra a las larvas y pupas de Glypholoma pustuliferum Jeannel recientemente colectadas en Chile y asignables a dicha especie por su asociaci&oacute;n con adultos y su correspondencia en tama&ntilde;o y forma entre pupas y adultos, ademas de sinapomorfias larvales conocidas en Staphylinidae, y conocimiento sobre otros elementos de la fauna chilena. Se realiz&oacute; un an&aacute;lisis filogen&#233;tico con base en car&aacute;cteres estructurales y quetot&aacute;xicos de estas larvas, adem&aacute;s de otras pertenecientes a 25 g&#233;neros del Grupo Omaliine. La posici&oacute;n de Glypholomatinae dentro del Grupo Omaliine permanece incierta aunque hay argumentos d&#233;biles de que forman un grupo monofil&#233;tico con Omaliinae y Microsilphinae. Se presentan argumentos que consolidan la monofilia de la subfamilia Proteininae y sus componentes y de la tribu Coryphiini de Omaliinae. 
ER  -

