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^A School of Land, Crop and Food Science, The University of Queensland, Gatton Campus, Gatton, Qld 4343, Australia.

^B School of Land, Crop and Food Science, The University of Queensland, Qld 4072, Australia.

^C Institut National de la Recherche Agronomique, Centre de Versailles-Grignon, Unité Environment et Grandes Cultures, 78850, Thierval-Grignon, France.

^D ARC Centre of Excellence for Integrative Legume Research, ARC Centre for Complex Systems and Advanced Computational Modelling Centre, The University of Queensland, Qld 4072, Australia.

^E Present address: Queensland Department of Primary Industries and Fisheries, Leslie Research Centre, Toowoomba, Qld 4350, Australia.

^F Corresponding author. Email: c.birch@uq.edu.au