Register      Login
The Rangeland Journal The Rangeland Journal Society
Journal of the Australian Rangeland Society
RESEARCH ARTICLE (Open Access)

Stocking density and distribution in relation to vegetation in a biodiverse semi-arid pastoral system in South Africa

M. Igshaan Samuels A B C * , Nicky Allsopp D and M. Timm Hoffman B
+ Author Affiliations
- Author Affiliations

A Agricultural Research Council – Animal Production, Biodiversity and Conservation Biology Department, University of the Western Cape, 7535, South Africa.

B Plant Conservation Unit, Department of Biological Sciences, University of Cape Town, 7701, South Africa.

C Biodiversity and Conservation Biology Department, University of the Western Cape, 7535, South Africa.

D South African Environmental Observation Network (SAEON), Fynbos Node, Claremont 7735, South Africa.

* Correspondence to: isamuels@uwc.ac.za

The Rangeland Journal 44(1) 1-12 https://doi.org/10.1071/RJ21046
Submitted: 18 August 2021  Accepted: 17 January 2022   Published: 7 March 2022

© 2022 The Author(s) (or their employer(s)). Published by CSIRO Publishing on behalf of the Australian Rangeland Society. This is an open access article distributed under the Creative Commons Attribution-NonCommercial-NoDerivatives 4.0 International License (CC BY-NC-ND)

Abstract

The spatial heterogeneity of vegetation in a communal grazing system provides pastoralists with a range of grazing sites to select for their livestock. When fencing spatially constrains herds, there is reduced access to ecological heterogeneity, which may have a negative impact on livestock and the grazing resource. This study investigated temporal and spatial changes in stocking densities and distribution within the six major vegetation types of the Leliefontein pastoral area in Namaqualand, South Africa. This semi-arid, biodiverse area of about 2000 km2 supports a spatially-constrained mobile pastoral system based on small stock production. Over the data period from 1999 to 2006, the location of 256 small stock herds, their temporal occupancy and herd size were used to determine stocking densities within the daily grazing radius of herds. These grazing radii were overlaid using GIS to produce stocking density distributions over the landscape. The density and location of herds were unevenly distributed when assessed monthly and annually. Parts of the study area were consistently overstocked whereas others remained ungrazed. We argue that the recommendation of fixed stocking rates within this variable environment is inappropriate because forage availability and quality fluctuate within and between years and within drought cycles. We conclude that development interventions should rather focus on reducing stocking densities during and after drought periods. This would benefit pastoralists through reduced livestock mortalities and the vegetation would recover more quickly from the impacts of drought.

Keywords: drought, grazing management, landscape ecology, livestock mobility, Namaqualand, overstocking, semiarid shrublands, spatial variation, stocking rate, Succulent Karoo.


References

Adler, PB, Raff, DA, and Lauenroth, WK (2001). The effect of grazing on the spatial heterogeneity of vegetation. Oecologia 128, 465–479.
The effect of grazing on the spatial heterogeneity of vegetation.Crossref | GoogleScholarGoogle Scholar |

Agrawal, A (1993). Mobility and cooperation among nomadic shepherds: the case of the Raikas. Human Ecology 21, 261–279.
Mobility and cooperation among nomadic shepherds: the case of the Raikas.Crossref | GoogleScholarGoogle Scholar |

Allen, VG, Batello, C, Berretta, EJ, Hodgson, J, Kothmann, M, Li, X, McIvor, J, Milne, J, Morris, C, Peeters, A, and Sanderson, M (2011). An international terminology for grazing lands and grazing animals. Grass and Forage Science 66, 2–28.
An international terminology for grazing lands and grazing animals.Crossref | GoogleScholarGoogle Scholar |

Anderson PML (2008) The impact of grazing along an environmental gradient in the Kamiesberg, South Africa. PhD Thesis, University of Cape Town, Cape Town, South Africa.

Anderson, GD, and Talbot, LM (1965). Soil factors affecting the distribution of the grassland types and their utilisation by wild animals on the Serengeti Plains, Tanganyika. Journal of Ecology 53, 33–56.
Soil factors affecting the distribution of the grassland types and their utilisation by wild animals on the Serengeti Plains, Tanganyika.Crossref | GoogleScholarGoogle Scholar |

Bailey, DW, Keil, MR, and Rittenhouse, LR (2004). Research observation: Daily movement patterns of hill climbing and bottom dwelling cows. Journal of Range Management 57, 20–28.
Research observation: Daily movement patterns of hill climbing and bottom dwelling cows.Crossref | GoogleScholarGoogle Scholar |

Bassett, TJ (1986). Fulani herd movements. Geographical Review 76, 233–248.
Fulani herd movements.Crossref | GoogleScholarGoogle Scholar |

Behnke RH, Scoones I (1993) Rethinking range ecology: implications for rangeland management in Africa. In ‘Range ecology at disequilibrium: new models of natural variability and pastoral adaptation in African savannas’. (Eds RH Behnke, I Scoones, C Kerven) pp. 1–30. (Overseas Development Institute: London, UK)

Beukes, PC, and Cowling, RM (2000). Impacts of non-selective grazing on cover, composition and productivity of Nama-karoo grassy shrubland. African Journal of Range and Forage Science 17, 27–35.

Campbell, BM, Gordon, IJ, Luckert, MK, Petheram, L, and Vetter, S (2006). In search of optimal stocking regimes in semi-arid grazing lands: one size does not fit all. Ecological Economics 60, 75–85.
In search of optimal stocking regimes in semi-arid grazing lands: one size does not fit all.Crossref | GoogleScholarGoogle Scholar |

Coppock, DL, Ellis, JE, and Swift, DM (1986). Livestock feeding ecology and resource utilization in a nomadic pastoral ecosystem. Journal of Applied Ecology 23, 573–583.
Livestock feeding ecology and resource utilization in a nomadic pastoral ecosystem.Crossref | GoogleScholarGoogle Scholar |

Davis, CL, MT Hoffman, M, and Roberts, W (2016). Recent trends in the climate of Namaqualand, a megadiverse arid region of South Africa. South African Journal of Science 112, 1–9.
Recent trends in the climate of Namaqualand, a megadiverse arid region of South Africa.Crossref | GoogleScholarGoogle Scholar |

DEA (Department of Environmental Affairs) (2013) ‘Long-Term Adaptation Scenarios Flagship Research Programme (LTAS) for South Africa.’ (Summary for Policy-Makers: Pretoria, South Africa)

Ellis, JE, and Swift, DM (1988). Stability of African pastoral ecosystems: alternate paradigms and implications for development. Journal of Range Management 41, 450–459.
Stability of African pastoral ecosystems: alternate paradigms and implications for development.Crossref | GoogleScholarGoogle Scholar |

Esler KJE, Milton SJ, Dean WRJ (Eds) (2006) ‘Karoo veld: ecology and management.’ (Briza Press: Pretoria, South Africa)

Fauchereau, N, Trzaska, S, Rouault, M, and Richard, Y (2003). Rainfall variability and changes in southern Africa during the 20th century in the global warming context. Natural Hazards 29, 139–154.
Rainfall variability and changes in southern Africa during the 20th century in the global warming context.Crossref | GoogleScholarGoogle Scholar |

Fritz, H, and Duncan, P (1994). On the carrying capacity for large ungulates of African Savanna Ecosystems. Biological Sciences 256, 77–82.
| 8008761PubMed |

Helme N, Desmet P (2006) ‘A description of the endemic flora and vegetation of the Kamiesberg uplands, Namaqualand, South Africa.’ (CEPF/SKEP: Cape Town, South Africa)

Hempson G (2010) Herbivore dynamics in an arid environment. PhD Thesis, University of Edinburgh, Edinburgh, Scotland.

Hendricks, HH, Midgley, JJ, Bond, WJ, and Novellie, PA (2004). Why communal pastoralists do what they do in the Richtersveld National Park. African Journal of Range and Forage Science 21, 29–36.
Why communal pastoralists do what they do in the Richtersveld National Park.Crossref | GoogleScholarGoogle Scholar |

Hendricks, HH, Clark, B, Bond, WJ, Midgley, JJ, and Novellie, PA (2005). Movement response patterns of livestock to rainfall variability in the Richtersveld National Park. African Journal of Range and Forage Science 22, 117–125.
Movement response patterns of livestock to rainfall variability in the Richtersveld National Park.Crossref | GoogleScholarGoogle Scholar |

Illius, AW, and O’Connor, TG (1999). On the relevance of nonequilibrium concepts to arid and semiarid grazing systems. Ecological Applications 9, 798–813.
On the relevance of nonequilibrium concepts to arid and semiarid grazing systems.Crossref | GoogleScholarGoogle Scholar |

Kotze D, Malan H, Ellery W, Samuels I, Saul L (2010). Assessment of the environmental condition, ecosystem service provision and sustainability of use of two wetlands in the Kamiesberg Uplands. Report No. TT439-09 Water Research Commission, Pretoria, 189 pp.

Low, WM, Dudzinski, ML, and Muller, WJ (1981). The influence of forage and climatic conditions on range community preference of shorthorn cattle in central Australia. Journal of Applied Ecology 18, 11–26.
The influence of forage and climatic conditions on range community preference of shorthorn cattle in central Australia.Crossref | GoogleScholarGoogle Scholar |

McCabe JT (1994) Mobility and land use among African pastoralists: Old conceptual problems and new interpretations. In ‘African pastoralist systems. An integrated approach’. (Eds E Fratkin, KA Galvin, EA Roth) pp. 69–90. (Lynne Rienner: London, UK)

Meissner HH (2000). Nutrient supplementation of the grazing animal. In ‘2000. Pasture management in South Africa’. (Ed. NM Tainton) p. 355. (University of Natal Press: Pietermaritzburg, South Africa)

Mucina L, Rutherford MC (2006) ‘The vegetation of South Africa, Lesotho and Swaziland’, (South African National Biodiversity Institute: Pretoria)

Niamir M (1991) ‘Traditional African range management techniques: implications for range development.’ (Paper 31d, Pastoral Development Network, Overseas Development Institute)

Niamir-Fuller M (1999) Conflict management and mobility among pastoralists in Karamoja, Uganda. In ‘Managing mobility in African rangelands: the legitimization of transhumance’. (Ed M Niamir-Fuller) pp. 149–183. (Intermediate Technology Publications: London)

Niamir-Fuller M, Turner MD (1999) A review of recent literature on pastoralism and transhumance in Africa. In ‘Managing mobility in African rangelands: the legitimization of transhumance’. (Ed. M Niamir-Fuller) pp. 18–46. (Intermediate Technology Publications: London)

O’Brien, EM (1993). Climatic gradients in woody plant species richness: towards an explanation based on an analysis of southern Africa’s woody flora. Journal of Biogeography 20, 181–198.

Riginos, C, and Hoffman, MT (2003). Changes in population biology of two succulent shrubs along a grazing gradient. Journal of Applied Ecology 40, 615–625.
Changes in population biology of two succulent shrubs along a grazing gradient.Crossref | GoogleScholarGoogle Scholar |

Samuels MI (2013) Pastoral mobility in a variable and spatially constrained South Africa environment. PhD Thesis, University of Cape Town, Cape Town, South Africa.

Samuels, MI, Allsopp, N, and Knight, RS (2007). Patterns of resource use by livestock during and after drought in a communal rangeland in Namaqualand, South Africa. Journal of Arid Environments 70, 728–739.
Patterns of resource use by livestock during and after drought in a communal rangeland in Namaqualand, South Africa.Crossref | GoogleScholarGoogle Scholar |

Samuels, MI, Allsopp, N, and Hoffman, MT (2008). Mobility patterns of livestock keepers in semi-arid communal rangelands of Namaqualand, South Africa. Nomadic Peoples 12, 123–148.
Mobility patterns of livestock keepers in semi-arid communal rangelands of Namaqualand, South Africa.Crossref | GoogleScholarGoogle Scholar |

Samuels, MI, Cupido, C, Swarts, MB, Palmer, AR, and Paulse, JW (2016). Feeding ecology of four livestock species under different management in a semi-arid pastoral system in South Africa. African Journal of Range and Forage Science 33, 1–9.
Feeding ecology of four livestock species under different management in a semi-arid pastoral system in South Africa.Crossref | GoogleScholarGoogle Scholar |

Samuels, MI, Swarts, MBV, Schroeder, A, Ntombela, KP, and Cupido, CF (2018). Through the lens of a herder: insights into landscape ethno-ecological knowledge on rangelands in Namaqualand. Anthropology Southern Africa 41, 136–152.
Through the lens of a herder: insights into landscape ethno-ecological knowledge on rangelands in Namaqualand.Crossref | GoogleScholarGoogle Scholar |

Samuels, MI, Allsopp, N, and Hoffman, MT (2019). Traditional mobile pastoralism in a contemporary semi-arid rangeland in Namaqualand, South Africa. Rangeland Ecology and Management 72, 195–203.
Traditional mobile pastoralism in a contemporary semi-arid rangeland in Namaqualand, South Africa.Crossref | GoogleScholarGoogle Scholar |

Schareika, N (2001). Environmental knowledge and pastoral migration among the Wodaabe of South-Eastern Niger. Nomadic Peoples 5, 61–88.
Environmental knowledge and pastoral migration among the Wodaabe of South-Eastern Niger.Crossref | GoogleScholarGoogle Scholar |

Scholes, RJ, Dowty, PR, Caylor, K, Parsons, DAB, Frost, PGH, and Shugart, HH (2009). Trends in savanna structure and composition along an aridity gradient in the Kalahari. Journal of Vegetation Science 13, 419–428.
Trends in savanna structure and composition along an aridity gradient in the Kalahari.Crossref | GoogleScholarGoogle Scholar |

Scholte P, Brouwer J (2008). Relevance of key resource areas for large-scale movements of livestock. In ‘Resource ecology: spatial and temporal dynamics of foraging’. (Eds HHT Prins, F van Langevelde) pp. 211–232. (Springer: The Netherlands)

Scoones, IE (1991). Wetlands in drylands: key resources for agricultural and pastoral production in Africa. Ambio 20, 366–371.

Scoones I (Ed) (1995) ‘Living with uncertainty: new directions in pastoral development in Africa.’ (Intermediate Technology Publications: London)

Sullivan, S, and Rohde, R (2002). On non-equilibrium in arid and semi-arid grazing systems. Journal of Biogeography 29, 1595–1618.
On non-equilibrium in arid and semi-arid grazing systems.Crossref | GoogleScholarGoogle Scholar |

Todd, SW (2006). Gradients in vegetation cover, structure and species richness of Nama-Karoo shrublands in relation to distance from livestock watering points. Journal of Applied Ecology 43, 293–304.
Gradients in vegetation cover, structure and species richness of Nama-Karoo shrublands in relation to distance from livestock watering points.Crossref | GoogleScholarGoogle Scholar |

Todd, SW, and Hoffman, MT (1999). A fence-line contrast reveals effect of heavy grazing on plant diversity and community composition in Namaqualand, South Africa. Plant Ecology 142, 169–17.
A fence-line contrast reveals effect of heavy grazing on plant diversity and community composition in Namaqualand, South Africa.Crossref | GoogleScholarGoogle Scholar |

Todd, SW, and Hoffman, MT (2009). A fence line in time demonstrates grazing-induced vegetation shifts and dynamics in the semiarid succulent Karoo. Ecological Applications 19, 1897–1908.
A fence line in time demonstrates grazing-induced vegetation shifts and dynamics in the semiarid succulent Karoo.Crossref | GoogleScholarGoogle Scholar | 19831078PubMed |

Valentine, JF (2000). ‘Grazing management’, 2nd edn. (Academic Press: San Diego, CA, USA)

Vetter, S (2005). Rangelands at equilibrium and non-equilibrium: recent developments in the debate around rangeland ecology and management. Journal of Arid Environments 62, 321–341.
Rangelands at equilibrium and non-equilibrium: recent developments in the debate around rangeland ecology and management.Crossref | GoogleScholarGoogle Scholar |