Invertebrate Systematics Invertebrate Systematics Society
Systematics, phylogeny and biogeography

Revision and cladistic analysis of the spider genus Carapoia González-Sponga (Araneae : Pholcidae), with descriptions of new species from Brazil’s Atlantic forest

Bernhard A. Huber
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- Author Affiliations

Alexander Koenig Zoological Research Museum, Adenauerallee 160, 53113 Bonn, Germany. Email:

Invertebrate Systematics 19(6) 541-556
Submitted: 16 August 2005  Accepted: 29 September 2005   Published: 16 January 2006


The pholcid genus Carapoia is revised and interspecific relationships are analysed cladistically. Five new species from the Brazilian Atlantic forest are described (C. ubatuba, C. brescoviti, C. una, C. crasto, C. rheimsae), and new records are given for the four previously described species. Cladistic analysis reveals a biogeographic split between a northern clade (Amazon, Venezuela, Guyana) and a southern clade (Brazilian Atlantic forest). While each of the three northern species is widely distributed, the six Atlantic forest species seem to be restricted to much smaller areas of forest remnants along the Atlantic coast.

Additional keywords: biogeography, morphology, phylogeny.


Material studied herein was kindly lent by Alexandre Bonaldo (MPEG), Antonio Brescovit and Cristina Rheims (IBSP) and Osvaldo Villarreal (MHNLS). I am deeply indebted to A. Brescovit and C. Rheims for their hospitality and support in Brazil in 2003 and to O. Villarreal and Abel Pérez González for their support in Venezuela in 2002. Two anonymous referees and A. Brescovit and C. Rheims commented on a previous version of the manuscript. Venezuelan material was collected under permit No 01–11–0966 issued by the Dirección General de Fauna y Oficina Nacional de Diversidad Biológica in Caracas. This work is part of BIOTA/FAPESP – The Biodiversity Virtual Institute Program.


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