International Journal of Wildland Fire International Journal of Wildland Fire Society
Journal of the International Association of Wildland Fire

Big fires and their ecological impacts in Australian savannas: size and frequency matters

Cameron P. Yates A B C , Andrew C. Edwards A B and Jeremy Russell-Smith A B

A Bushfires NT, PO Box 37346, Winnellie, NT 0821, Australia.

B Tropical Savannas Cooperative Research Centre, Charles Darwin University, Building 31, Level 3, Darwin, NT 0909, Australia.

C Corresponding author. Email:

International Journal of Wildland Fire 17(6) 768-781
Submitted: 15 October 2007  Accepted: 13 May 2008   Published: 12 December 2008


Savannas are the most fire-prone of the earth’s major biomes. The availability of various broad-scale satellite-derived fire mapping and regional datasets provides a framework with which to examine the seasonality, extent and implications of large fires with particular reference to biodiversity values in the tropical savannas of northern Australia. We document the significance of savanna fires in the fire-prone ‘Top End’ region of the Northern Territory, Australia, using 9 years (1997–2005) of National Oceanic and Atmospheric Administration (NOAA)-Advanced Very High Resolution Radiometer (AVHRR)-, Landsat Thematic Mapper (TM)- and Enhanced Thematic Mapper (ETM+)-derived fire mapping. Fire (patch) sizes from both AVHRR- and Landsat-scale mapping increased through the calendar year associated with progressive curing of grass and litter fuels. Fire frequency data at both satellite sensor scales indicate that regional fire regimes in higher rainfall regions are dominated by large (>1000 km2) fires occurring typically at short (~2–3 years) fire return intervals. In discussion, we consider the ecological implications of these patch size distributions on regional fire-sensitive biota. Collectively, assembled data illustrate that many northern Australian savanna flora, fauna and habitats embedded within the savanna matrix are vulnerable to extensive and frequent fires, especially longer-lived obligate seeder plant taxa and relatively immobile vertebrate fauna with small home ranges.

Additional keywords: Arnhem Land, AVHRR, fire-affected area, Landsat, patch size, Top End.


AGO (2007) ‘National Greenhouse Gas Inventory 2006.’ (Australian Greenhouse Office: Canberra)

Andersen AN, Cook GD, Williams RJ (Eds) (2003) ‘Fire in Tropical Savannas: the Kapalga Experiment.’ (Springer-Verlag: New York)

Andreae MO (1997) Emissions of trace gases and aerosols from southern African fires. In ‘Fires in Southern African Savannas; Ecological and Atmospheric Perspectives’. (Eds BW van Wilgen, MO Andreae, JG Goldammer, JA Lindsay) pp. 161–184. (Witwatersrand University Press: Johannesburg)

Barrow PH (2008) The role of fire in the ecology of Leichhardt’s grasshopper (Petasida ephippigera) and its food plants, Pityrodia spp. PhD thesis, Charles Darwin University, Darwin.

Barrow PHDuff GALiddle DRussell-Smith J1993Threats to monsoon rainforest habitat in northern Australia: the case of Ptychosperma bleeseri Burrett (Arecaceae).Australian Journal of Ecology18463471doi:10.1111/J.1442-9993.1993.TB00472.X

Begg RJ1981The mammals of Little Nourlangie rock, NT. IV. Ecology of Zyzomys woodwardi, the large rock-rat, and Zyzomys argurus, the common rock-rat (Rodentia: Muridae).Australian Wildlife Research8307320doi:10.1071/WR9810307

Beringer JPackham DTapper N1995Biomass burning and resulting emissions in the Northern Territory, Australia.International Journal of Wildland Fire5229235doi:10.1071/WF9950229

Blake G (2004) Object-oriented mapping of sandstone heath vegetation on the Arnhem Plateau. BSc(Hons) thesis, Charles Darwin University, Darwin.

Bowman DMJS1991Environmental determinants of Allosyncarpia ternata forests that are endemic to western Arnhem Land, northern Australia.Australian Journal of Botany39575589doi:10.1071/BT9910575

Bowman DMJS1994Preliminary observations on the mortality of Allosyncarpia ternata stems on the Arnhem Land Plateau, northern Australia.Australian Forestry576264

Bowman DMJS1997Observations on the demography of the Australian boab (Adansonia gibbosa) in the north-west of the Northern Territory, Australia.Australian Journal of Botany45893904

Bowman DMJSPanton WJ1993Decline of Callitris intratropica in the Northern Territory: implications for pre- and post-colonisation fire regimes.Journal of Biogeography20373381doi:10.2307/2845586

Bowman DMJSWilson BAHooper RJ1988Response of Eucalyptus forest and woodland to four fire regimes at Munmarlary, Northern Territory.Australian Journal of Ecology76215232

Braithwaite RW, Estbergs JA (1985) Fire patterns and woody vegetation trends in the Alligator Rivers Region of northern Australia. In ‘Ecology and Management of the World’s Savannas’. (Eds JC Tothill, JJ Mott) pp. 359–364. (Australian Academy of Science: Canberra)

Braithwaite RWGriffiths AD1994Demographic variation and home range contraction in the northern quoll, Dasyurus hallaculatus (Marsupialia: Dasyuridae).Wildlife Research21203217

Brockett BHBiggs HCvan Wilgen BW2001A patch mosaic burning system for conservation areas in southern African savannas.International Journal of Wildland Fire10169183doi:10.1071/WF01024

Burbidge AAMcKenzie NL1989Patterns in the modern decline of Western Australia’s vertebrate fauna; causes and conservation implications.Biological Conservation50143198doi:10.1016/0006-3207(89)90009-8

Cook GDHurst DFGriffith DWT1995Atmospheric trace gas emissions from tropical Australian savanna fires.CALMscience4Suppl.123128

Craig R, Heath B, Raisbeck-Brown N, Steber M, Marsden J, Smith R (2002) The distribution, extent and seasonality of large fires in Australia, April 1998–March 2000, as mapped from NOAA-AVHRR imagery. In ‘Australian Fire Regimes: Contemporary Patterns (April 1998–March 2000) and Changes since European Settlement’, Australia: State of the Environment Second Technical Paper Series (Biodiversity), Series 2. (Department of Environment and Heritage: Canberra) Available at [Verified 3 December 2008]

Crisp MDLaffan SLinder HPMonro A2001Endemism in the Australian flora.Journal of Biogeography28183198doi:10.1046/J.1365-2699.2001.00524.X

Csiszar IDenis LGiglio LJustice COHewson J2005Global fire activity from two years of MODIS data.International Journal of Wildland Fire14117130doi:10.1071/WF03078

Dostine PLJohnson GCFranklin DCZhang YHempel C2001Seasonal use of savanna landscapes by the Gouldian Finch, Erythrura gouldiae, in the Yinberrie Hills area, Northern Territory.Wildlife Research28445458doi:10.1071/WR00049

Dwyer EPinnock SGregoire J-MPereira JMC2000Global spatial and temporal distribution of vegetation fire as determined from satellite observations.International Journal of Remote Sensing2112891302doi:10.1080/014311600210182

Dyer R, Jacklyn P, Partridge I, Russell-Smith J, Williams RJ (Eds) (2003) ‘Savanna Burning: Understanding and Using Fire in Northern Australia.’ (Tropical Savannas Cooperative Research Centre: Darwin)

Edwards AC, Russell-Smith J, Ecological thresholds and the status of fire-sensitive vegetation in western Arnhem Land, northern Australia. International Journal of Wildland Fire, in press. doi:10.1071/WF08008

Edwards ACHauser PAnderson MMcCartney JArmstrong MThackway RAllan GEHempel CRussell-Smith J2001A tale of two parks: contemporary fire regimes of Litchfield and Nitmiluk National Parks, monsoonal northern Australia.International Journal of Wildland Fire107989doi:10.1071/WF01002

Ellis S, Kanowski P, Whelan R (2004) Report of the National Inquiry on Bushfire Mitigation and Management. (Commonwealth of Australia: Canberra) Available at [Verified 21 November 2008]

Felderhof L (2007) The fire patchiness paradigm: a case study in northwest Queensland. PhD thesis, James Cook University, Cairns.

Felderhof LGillieson D2006Comparison of fire patterns and fire frequency in two tropical savanna biomes.Austral Ecology31736746doi:10.1111/J.1442-9993.2006.01645.X

Firth RSCWoinarski JCZNoske RA2006Home range and den characteristics of the brush-tailed rabbit-rat (Conillurus penicillatus) in the monsoonal tropics of the Northern Territory.Australian Wildlife Research33397408doi:10.1071/WR05057

Fisher RVigilante TYates CPRussell-Smith J2003Patterns of landscape fire and predicted vegetation response in the North Kimberley region of Western Australia.International Journal of Wildland Fire12369379doi:10.1071/WF03021

Franklin DC1999Evidence of disarray amongst granivorous bird assemblages in the savannas of northern Australia, a region of sparse human settlement.Biological Conservation905368doi:10.1016/S0006-3207(99)00010-5

Franklin DCWhitehead PJPardon GMatthews JMcMahon PMcIntyre D2005Geographic patterns and correlates of the decline of granivorous birds in northern Australia.Wildlife Research32399408doi:10.1071/WR05052

Fraser FJ (2000) The impacts of fire and grazing on the partridge pigeon: the ecological requirements of a declining tropical granivore. PhD thesis, Australian National University, Canberra.

Fraser FJLawson VMorrison SChristopherson PMcGregor SRawlinson M2003Fire management experiment for the declining partridge pigeon, Kakadu National Park.Ecological Management & Restoration494102doi:10.1046/J.1442-8903.2003.00142.X

Friend GRTaylor JA1985Habitat preferences of small mammals in tropical open-forests of northern Australia.Australian Journal of Ecology10173185doi:10.1111/J.1442-9993.1985.TB00879.X

Gill AMAllan GEYates CP2003Fire-created patchiness in Australian savannas.International Journal of Wildland Fire12323331doi:10.1071/WF03018

Graham G (2002) An assessment of the distribution and status of Arnhem cypress pine Callitris intratropica (R. T. Baker & H. G. Sm.) in the Kimberley Region, Western Australia. Western Australia Department of Conservation and Land Management, report to the National Reserves System Program, Environment Australia. (Perth)

Griffiths ADKoenig JCarrol FPrice O2002Activity area and day-time tree use of the black-footed tree-rat Mesembriomys gouldii.Australian Mammalogy23181183

Ingwersen F (1995) Kakadu–Alligator Rivers Region, Northern Territory, Australia—CPD Site Au4. In ‘Centres of Plant Diversity: a Guide and Strategy for their Conservation. Vol. 2. Asia, Australasia and the Pacific’. (Eds SD Davis, VH Heywood, AC Hamilton) pp. 471–475. (World Wildlife Fund and International Union for Conservation of Nature: Cambridge, UK)

Kasischke KSFrench NHF1995Locating and estimating the aerial extent of wildfires in Alaskan boreal forests using multiple-season AVHRR NDVI composite data.Remote Sensing of Environment51263275

Keeley JKFotheringham CJMorais M1999Re-examining fire suppression impacts on brushland fire regimes.Science28418291832doi:10.1126/SCIENCE.284.5421.1829

Keith DA, McCaw WL, Whelan RJ (2002a) Fire regimes in Australian heathlands: fire fuels, behaviour and effects on plants and animals. In ‘Flammable Australia: the Fire Regimes and Biodiversity of a Continent’. (Eds RA Bradstock, JE Williams, AM Gill) pp. 197–237. (Cambridge University Press: Cambridge, UK)

Keith DA, Williams JE, Woinarski JCZ (2002b) Fire management and biodiversity conservation: key approaches and principles. In ‘Flammable Australia: the Fire Regimes and Biodiversity of a Continent’. (Eds RA Bradstock, JE Williams, AM Gill) pp. 401–425. (Cambridge University Press: Cambridge, UK)

Kerle JA1998The population dynamics of a tropical possum, Trichosurus vulpecula arnhemensis Collett.Wildlife Research25171182doi:10.1071/WR96113

Lee AK, Copley P, Morris K, Calaby JH, Flannery TF (1995) ‘The Action Plan for Australian Rodents.’ (Department Environment and Heritage: Canberra)

Liddle DT (2004) The ecology of Cycas armstrongii and management of fire in Australia’s tropical savannas. PhD thesis, Charles Darwin University, Darwin.

Liddle DTBrook BWMatthews JTaylor SMCaley P2006Treat and response: a decade of decline in a regionally endangered rainforest palm affected by fire and introduced animals.Biological Conservation132362375doi:10.1016/J.BIOCON.2006.04.028

Lonsdale WMBraithwaite RW1991Assessing the effects of fire on vegetation in tropical savannas.Australian Journal of Ecology16363374doi:10.1111/J.1442-9993.1991.TB01064.X

Lowe L1995Preliminary investigations of the biology and management of Leichhardt’s grasshopper, Petasida ephippigera White.Journal of Orthoptera Research4219221doi:10.2307/3503479

Luke RH, McArthur AG (1978) ‘Bushfires in Australia.’ (Australian Government Publishing Service: Canberra)

Maxwell S, Burbidge A, Morris K (1996) ‘The 1996 action plan for Australian marsupials and monotremes. IUCN/SSC Australian Marsupial and Monotreme Specialist Group.’ (Wildlife Australia: Canberra)

Noske R1992The status and ecology of the white-throated grasswren Amytornis woodwardi.Emu923951

Palmer CPrice OBach C2000Foraging ecology of the black flying fox (Pteropus alecto) in the seasonal tropics of the Northern Territory.Australian Wildlife Research27169178

Pardon LGBrook BWGriffiths ADBraithwaite RW2003Determinants of survival for the northern brown bandicoot under a landscape-scale fire experiment.Journal of Animal Ecology72106115doi:10.1046/J.1365-2656.2003.00686.X

Parr CAndersen AN2006Patch mosaic burning for biodiversity conservation: a critique of the pyrodiversity paradigm.Conservation Biology2016101619doi:10.1111/J.1523-1739.2006.00492.X

Pausas JGBradstock RAKeith DAKeeley JE2004Plant functional traits in relation to fire in crown-fire ecosystems.Ecology8510851100doi:10.1890/02-4094

Perry J (2005) Status and habitat characteristics of the endemic Carpentarian grasswren (Amytornis dorotheae) in the Northern Territory. MSc thesis, Charles Darwin University, Darwin.

Petty AMWerner PALehmann CERRiley JEBanfai DSElliott LP2007Savanna responses to feral buffalo in Kakadu National Park, Australia.Ecological Monographs77441463doi:10.1890/06-1599.1

Preece N2007Traditional and ecological fires and effects of bushfire laws in north Australian savannas.International Journal of Wildland Fire16378389doi:10.1071/WF05079

Press AJ1988Comparisons of the extent of fire in different land management systems in the Top End of the Northern Territory.Proceedings of the Ecological Society of Australia15167175

Price ORussell-Smith JEdwards A2003Fine-scale patchiness of different fire intensities in sandstone heath vegetation in northern Australia.International Journal of Wildland Fire12227236

Price ORankmore BMilne DBrock CTynan CKean LRoeger L2005Regional patterns of mammal abundance and their relationship to landscape variables in eucalypt woodlands near Darwin.Wildlife Research32435446doi:10.1071/WR04033

Price OEdwards ARussell-Smith J2007The efficacy of permanent firebreaks and aerial prescribed burning in western Arnhem Land, Northern Territory, Australia.International Journal of Wildland Fire16295307doi:10.1071/WF06039

Price OF2006Movements of frugivorous birds among fragmented rainforests in the Northern Territory.Australian Wildlife Research33521528doi:10.1071/WR06029

Price OFWoinarski JCZRobinson D1999Very large area requirements for frugivorous birds in monsoon rainforests of the Northern Territory, Australia.Biological Conservation91169180doi:10.1016/S0006-3207(99)00081-6

Prior LDBrook BWWilliams RJWerner PABradshaw CJABowman DMJS2006Environmental and allometric drivers of tree growth rates in a north Australian savanna.Forest Ecology and Management234164180doi:10.1016/J.FORECO.2006.06.034

Puckey HLewis MHooper DMichell C2004Home range, movement and habitat utilisation of the Carpentarian rock-rat (Zyzomys palatalis) in an isolated habitat patch.Wildlife Research31327337doi:10.1071/WR03025

Rankmore BR (2006) Impacts of habitat fragmentation on the vertebrate fauna of the tropical savannas of Northern Australia, with special reference to medium-sized mammals. MSc thesis, Charles Darwin University, Darwin.

Roy DPJin YLewis PEJustice CO2005Prototyping a global algorithm for systematic fire-affected area mapping using MODIS time series data.Remote Sensing of Environment97137162doi:10.1016/J.RSE.2005.04.007

Russell-Smith J2006Recruitment dynamics of the long-lived obligate seeders Callitris intratropica (Cupressaceae) and Petraeomyrtus punicea (Myrtaceae).Australian Journal of Botany54479485doi:10.1071/BT05133

Russell-Smith JEdwards AC2006Seasonality and fire severity in savanna landscapes of monsoonal northern Australia.International Journal of Wildland Fire15541550doi:10.1071/WF05111

Russell-Smith JLee AH1992Plant populations and monsoon rain forest in the Northern Territory, Australia.Biotropica24471487doi:10.2307/2389010

Russell-Smith JSetterfield SA2006Monsoon rain forest seedling dynamics, northern Australia: contrasts with regeneration in eucalypt-dominated savannas.Journal of Biogeography3315971614doi:10.1111/J.1365-2699.2006.01527.X

Russell-Smith J, Stanton JP (2002) Fire regimes and fire management of rainforest communities across northern Australia. In ‘Flammable Australia’. (Eds RA Bradstock, JE Williams, AM Gill) pp. 329–350. (Cambridge University Press: Cambridge, UK)

Russell-Smith JYates CP2007Australian savanna fire regimes: context, scale, patchiness.Fire Ecology34863

Russell-Smith J, McKenzie NL, Woinarski JCZ (1992) Conserving vulnerable habitat in northern and north-western Australia: the rainforest archipelago. In ‘Conservation and Development Issues in Northern Australia’. (Eds I Moffatt, A Webb) pp. 64–69. (North Australia Research Unit of the Australian National University: Darwin)

Russell-Smith JLucas DEBrock JBowman DMJS1993Allosyncarpia-dominated rain forest in monsoonal northern Australia.Journal of Vegetation Science46782

Russell-Smith JRyan PGDuRieu R1997A LANDSAT MSS-derived fire history of Kakadu National Park, monsoonal northern Australia, 1980–1994: seasonal extent, frequency and patchiness.Journal of Applied Ecology34748766doi:10.2307/2404920

Russell-Smith JRyan PGKlessa DWaight GHarwood R1998Fire regimes, fire-sensitive vegetation, and fire management of the sandstone Arnhem Plateau, monsoonal northern Australia.Journal of Applied Ecology35829846doi:10.1046/J.1365-2664.1998.3540553.X

Russell-Smith JRyan PGCheal D2002bFire regimes and the conservation of sandstone heath in monsoonal northern Australia: frequency, interval, patchiness.Biological Conservation10491106doi:10.1016/S0006-3207(01)00157-4

Russell-Smith JEdwards ACook GD2003aReliability of biomass burning estimates from savanna fires: biomass burning in northern Australia during the 1999 Biomass Burning and Lightning Experiment-B field campaign.Journal of Geophysical Research – Atmospheres108D38405doi:10.1029/2001JD000787

Russell-Smith JYates CPEdwards AAllan GECook GDCooke PCraig RHeath BSmith R2003bContemporary fire regimes of northern Australia: change since Aboriginal occupancy, challenges for sustainable management.International Journal of Wildland Fire12283297doi:10.1071/WF03015

Russell-Smith JWhitehead PJCook GDHoare JL2003cResponse of Eucalyptus-dominated savanna to frequent fires: lessons from Munmarlary, 1973–1996.Ecological Monographs73349375doi:10.1890/01-4021

Russell-Smith JYates CPLynch B2006Fire regimes and soil erosion in north Australian hilly savannas.International Journal of Wildland Fire15551556doi:10.1071/WF05112

Russell-Smith JYates CPWhitehead PJSmith RCraig RAllan GThackway RFrakes ICridland SMeyer MCPGill AM2007Bushfires ‘Down Under’: patterns and implications of Australian landscape burning.International Journal of Wildland Fire16361377doi:10.1071/WF07018

Shapcott A2000Conservation and genetics in the fragmented monsoon rainforest in the Northern Territory, Australia: a case study of three frugivorous dispersed species.Australian Journal of Botany48397407doi:10.1071/BT98081

Stocks BJMason JATodd JBBosch EMWotton BMAmiro BDFlannigan MDHirsch KGLogan KAMartell DLSkinner WR2003Large forest fires in Canada, 1989–1997.Journal of Geophysical Research1078149doi:10.1029/2001JD000484

Strahan R (Ed.) (1995) ‘The Mammals of Australia.’ (Reed Books: Sydney)

Stocker GC (1966) Aspects of the seeding habits of Callitris intratropica. Commonwealth of Australia, Department of National Development, Forestry and Timber Bureau, Leaflet no. 97. (Canberra)

Telfer WRGriffiths AD2006Dry-season use of space, habitats and shelters by the short-eared rock-wallaby (Petrogale brachyotis) in the monsoon tropics.Wildlife Research33207214doi:10.1071/WR05032

Thackway R, Cresswell I (1995) ‘An Interim Biogeographic Regionalisation for Australia: a Framework for Establishing the National System of Reserves, Version 4.0.’ (Australian Nature Conservation Agency: Canberra)

Townsend SADouglas MM2000The effect of three fire regimes on stream water quality, water yield and export co-efficients in a tropical savanna (northern Australia).Journal of Hydrology229118137doi:10.1016/S0022-1694(00)00165-7

Trainor CFisher AWoinarski JChurchill S2000Multiscale patterns of habitat use by the Carpentarian rock-rat (Zyzomys palatalis) and the common rock-rat (Z. argurus).Wildlife Research27319332doi:10.1071/WR97040

Walker J (1981) Fuel dynamics in Australian vegetation. In ‘Fire and the Australian Biota’. (Eds AM Gill, RM Groves, IR Noble) pp. 101–127. (Australian Academy of Science: Canberra)

Werner PA2005Impact of feral water buffalo and fire on growth and survival of mature savanna trees: an experimental field study in Kakadu National Park, northern Australia.Austral Ecology30625647doi:10.1111/J.1442-9993.2005.01491.X

Werner PACowie IDCusack JS2006Impact of feral water buffalo on juvenile trees in savannas of northern Australia: an experimental field study in Kakadu National Park.Australian Journal of Botany54283296doi:10.1071/BT04135

Williams RJCook GDGill AMMoore PHR1999Fire regime, fire intensity and tree survival in a tropical savanna in northern Australia.Australian Journal of Ecology245059doi:10.1046/J.1442-9993.1999.00946.X

Williams RJ, Griffiths AD, Allan GE (2002) Fire regimes and biodiversity in the wet-dry tropical landscapes of northern Australia. In ‘Flammable Australia: the Fire Regimes and Biodiversity of a Continent’. (Eds RA Bradstock, JE Williams, AM Gill) pp. 281–304. (Cambridge University Press: Cambridge, UK)

Williams RJ, Gill AM, Moore PHR (2003) Fire behaviour. In ‘Fire in Tropical Savannas: the Kapalga Experiment’. (Eds AN Andersen, GD Cook, RJ Williams) pp. 33–46. (Springer-Verlag: New York)

Woinarski JCZ1990Effects of fire on the bird communities of tropical woodlands and open forests in northern Australia.Australian Journal of Ecology15122doi:10.1111/J.1442-9993.1990.TB01016.X

Woinarski JCZ1992The conservation status of the white-throated grasswren Amytornis woodwardi, an example of problems in status designation.Northern Territory Naturalist1315

Woinarski JCZFisher A1995Wildlife of lancewood (Acacia shirleyi) thickets and woodlands in northern Australia. 1. Variation in vertebrate species composition across the environmental range occupied by lancewood vegetation in the Northern Territory.Wildlife Research22379411

Woinarski JCZTidemann SC1991The bird fauna of a deciduous woodland in the wet–dry tropics of northern Australia.Wildlife Research18479500doi:10.1071/WR9910479

Woinarski JCZBrock CFisher AMilne DOliver B1999Response of birds and reptiles to fire regimes on pastoral lands in the Victoria River District, Northern Territory.The Rangeland Journal212438doi:10.1071/RJ9990024

Woinarski JCZBrock CArmstrong MHempel CCheal DBrennan K2000Bird distribution in riparian vegetation in the extensive natural landscapes of Australia’s tropical savanna: a broad-scale survey and analysis of a distributional data base.Journal of Biogeography27843868doi:10.1046/J.1365-2699.2000.00439.X

Woinarski JCZMilne DWanganeen G2001Changes in mammal populations in relatively intact landscapes of Kakadu National Park, Northern Territory, Australia.Austral Ecology26360370doi:10.1046/J.1442-9993.2001.01121.X

Woinarski JCZRisler JKean L2004The response of vegetation and vertebrate fauna to 23 years of fire exclusion in a tropical eucalyptus open forest, Northern Territory, Australia.Austral Ecology29156176doi:10.1111/J.1442-9993.2004.01333.X

Woinarski JCZWilliams RJPrice ORankmore B2005Landscapes without boundaries: wildlife and their environments in northern Australia.Wildlife Research32377388doi:10.1071/WR03008

Woinarski JCZHempel CCowie IBrennan KKerrigan RLeach GRussell-Smith J2006Distributional pattern of plant species restricted to the Northern Territory, Australia.Australian Journal of Botany54627640doi:10.1071/BT05041

WWF (2005) Nomination for listing Arnhem Plateau Sandstone Heath as an Endangered Ecological Community under Australia’s Environment Protection and Biodiversity Conservation Act 1999. Unpublished document lodged with the Commonwealth of Australia (Department of Environment Water Heritage and the Arts), Canberra. (WWF Australia: Darwin)

Yates CP, Russell-Smith J (2002) An assessment of the accuracy of DOLA’s northern Australian NOAA-AVHRR fire-affected area (FAA) map products. In ‘Australian Fire Regimes: Contemporary Patterns (April 1998–March 2000) and Changes since European Settlement’, Australia: State of the Environment Second Technical Paper Series No. 2 (Biodiversity). (Eds J Russell-Smith, R Craig, AM Gill, R Smith, JE Williams) (Department of the Environment and Heritage: Canberra) Available at [Verified 3 December 2008]

Yates CPRussell-Smith J2003Fire regimes and vegetation sensitivity analysis: an example from Bradshaw Station, monsoonal northern Australia.International Journal of Wildland Fire12349358doi:10.1071/WF03019

Yibarbuk DWhitehead PJRussell-Smith JJackson DGodjuwa CCooke PChoquenot DBowman DMJS2001Fire ecology and Aboriginal land management in central Arnhem Land, northern Australia: a tradition of ecosystem management.Journal of Biogeography28325344doi:10.1046/J.1365-2699.2001.00555.X

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